Polydrusus (Conocetus) angustus ( Lucas, 1854 )

Christoph Germann, 2018, A review of Conocetus Desbrochers des Loges, 1875, subgenus of Polydrusus Germar, 1817 (Coleoptera, Curculionidae, Entiminae), European Journal of Taxonomy 392, pp. 1-39 : 16-18

Polydrusus (Conocetus) angustus ( Lucas, 1854)

Figs 3C–D , 7F , 9H , 11

Material examined

Syntypes

Two syntypes of Polydrusus virens ( Kiesenwetter, 1864) :

GREECE: 1 ♀, Athens // Kiesenwetter (ZSM); 1 ♂, virens Ksw. Grèce Ksw. // Celadonius (Phyll.) Brull. ♂ Polydrosus [sic!] (ZSM). [Both with additional red labels:] Syntype Polydrusus (Conocetus) virens ( Kiesenwetter, 1864) labelled C. Germann 2016.

Two syntypes of Polydrusus (Conocetus) femoratus ( Stierlin, 1888) :

GREECE: 1 ♂, Griechenland [handwritten by L. Dieckmann] // Syntypus [red label] // coll. Stierlin // Polydrusus femoratus Stierl. det. Dieckmann 1989 (SDEI); 1 ♂, Griechenld // P. femoratus Stl. [both Stierlin’s handwriting] // Syntypus // Polydrusus femoratus Stierl. det. Dieckmann 1989 (SDEI).

Type locality

GREECE: Mount Ida, Crete.

Other material examined

GREECE: 1 ♀, Naxos, coll. Stierlin (SDEI); 4 ex., Graecia coll. Stierlin (SDEI); 1 ♀, Thessalia, Volo, leg. Stussiner (SDEI); 3 ex., Thessal[ia] Pelion, leg. Stussiner (SDEI); 2 ex., Aegina v. Oertzen (SDEI); 1 ♂, Naxos, leg. Schatzmayr (SDEI); 2 ex., Kreta, leg. Paganetti (SDEI); 7 ex., Parnass, leg. Paganetti (SDEI); 1 ex., Attica, leg. Reitter (SDEI); 1 ♂, Athen, coll. V. Heyden (SDEI); 1 ♀, Attica, leg. Reitter, coll. V. Heyden (SDEI); 1 ♂, Graecia leg. Kraatz (SDEI); 1 ♂, 1 ♀, Graecia, leg. Faust (SDEI); 2 ♀♀, Parnass, leg. Paganetti, coll. Piesbergen (SDEI); 2 ex., Asiat. Türkei, Rhodes, coll. Stöcklein (NMB); 2 ex., Volo, Thessalien, coll. P. Meyer (NMB); 2 ♂♂, Parnass, Graecia (NMB); 2 ex., Cycla. [Cyclades], coll. Kraatz (SDEI); 1 ex., femoratus Strl. Graecia Zeiller (SDEI); 1 ex., Taygetos, coll. Apfelbeck (NMB); 1 ex., Graecia, coll. Apfelbeck (NMB); 2 ex., Parnass, leg. Paganetti, coll. J. Breit (NMB); 3 ex., Parnass, leg. Paganetti 1903, coll. Stöcklein (NMB); 2 ex., Kreta, leg. Paganetti, coll. Stöcklein (NMB); 1 ex., Parnass, Paganetti, coll. G.A.K. Marshall (BMNH); 1 ♂, Parnass, Paganetti, coll. J.B. Jörger (NMB); 2 ex., Parnass, Paganetti, coll. J. Breit (NMB); 1 ex., Kreta, Paganetti, coll. G.A.K. Marshall (BMNH); 1 ex., Paleohoka, 17 May 1999, Crète, C. Besnard rec. (cJP); 4 ex., Saloniki, coll. A. Rätzer (NMBE); 1 ex., Kreta Ins., Dikti Mts, Lesithi plateau, 800 m, 5 Jun. 2001, leg. J. Voříšek (BMNH); 2 ex., Rhodos, 11–30 May 1974, v. Budberg leg. (BMNH); 1 ♂, Peloponesos, S of Tripolis, Karies, 24 Jun. 2004, J. Voříšek leg. (BMNH); 1 ♀, Peloponesos or., Astros, 5–20 m, 25 May 2004, J. Voříšek leg. (BMNH); 1 ex., Rhodos, 9 May [19]70, leg. Dr. Engelman, coll O. Voříšek (BMNH); 2 ♂, Attika, Varkiza, 6 May [19]93, leg. Colonnelli (cRB); 21 ex., Rhodes, Lachania, env. 50 m, 8 May 1995, leg. R. Borovec (cCG, cRB); 1 ex., Rhodes (FO 13), Afandou beach, 5 May [20]14, leg. J. Messutat (cJM); 18 ex., Rhodes, Kalithea env. (FO 1), 24 Apr.–7 May 2014, leg. J. Messutat (cJM); 4 ex., Rhodes (FO 2), Kolymbia, beach, 26 Apr. [20]14, leg. J. Messutat (cJM); 8 ex., Rhodes (FO 7), Apollona, N, 400 m, 29 Apr. [20]14, leg. J. Messutat (cJM); 1 ex., Rhodes (FO 3), Archipoli NE, 150 m, 26 Apr. [20]14, leg. J. Messutat (cJM); 1 ex., Rhodes (FO 6), Laerma N, 260 m, 28 Apr. [20]14, leg. J. Messutat (cJM); 7 ex., Rhodes (FO 9), Soroni-Fanes, 30 m, 1 May [20]14, leg. J. Messutat (cJM); 1 ex., Rhodes (FO 12), Genadi, beach, 3 May [20]14, leg. J. Messutat (cJM); Rhodes (FO 11), Emponas, 380 m, 3 May [20]14, leg. J. Messutat (cJM); 3 ex., Rhodes (FO 13), Afandou, beach, 5 May [20]14, leg. J. Messutat (cJM); 1 ♂, 1 ♀, Agios Nikolao, Kreta, 24 Apr. 1971, W. Wittmer (NMB); 1 ex., Crete, Chania, Potamida, 25 Apr. 2011, leg. H. Winkelmann (cHW); 2 ♂♂, 2 ♀♀, Attika, Sounio (SE Athens), 37°39′00″ N, 24°01′40″ E, 50 m, 17 May 2011, leg. Bahr, Bayer, Brunner and Winkelmann (cHW); 1 ex., Peloponnes, Kaliani, Prov. Korinthia, 5 May 2010, leg. W. Ziegler (cHW); 1 ex., Prov. Drama, Prosotsani, 20 Jul. 2009, leg. W. Ziegler (cHW); 1 ex., Prov. Korinthia, Stimfalia, 13 May 2007, leg. W. Ziegler (cHW); 1 ♂, Rhodos, Kallithea, 5–19 May 2005, leg. S. Scharf (cHW); 2 ♀♀, Kreta, Knossos Umgb., Jun. [19]92, leg. H.-J. Grunwald (cHW); 3 ex., Platistomo, 31 May 2013, 38°58′ N, 22°07′ E, leg. Eifler (cHW); 1 ♂, Peloponnese, Argolis, Mt Didymo, Didyma, doline, 6 Apr. 2010, leg. H. Winkelmann (cHW); 1 ♀, Peloponnese, Argolis, Palea Epidavros, 1–10 Apr. 2010, leg. H. Winkelmann (cHW); 1 ♀, Peloponnese, Corinthia, Korfos env., 10 Apr. 2010, leg. H. Winkelmann (cHW); 2 ♂, Crete, Chania, Potamida, 25 Apr. 2011, leg. H. Winkelmann (cHW); 1 ♀, Peloponnes, Galatas, 7–28 Jun. [19]76, Köstlin leg. (SDEI); 1 ♀, Samos, May [19]87, leg. Zimmerman (SDEI); 18 ex., Rhodes, Kalithea env. (FO1), 24 Apr.–7 May 2014, leg. Messutat (cCG, cJM, NMBE); 1 ♂, Rhodes (FO 12), Genadi, beach, 3 May [20]14, leg. Messutat (cJM); 2 ♂♂, 2 ♀♀, Rhodes (FO 13), Afandou, beach, 5 May [20]14, leg. Messutat (cJM); 1 ♀, Rhodes (FO 11), Emponas, 380 m, 3 May [20]14, leg. Messutat (cJM); 8 ex., Rhodes (FO 7),Apollona N, 400 m, 29 Apr.[20]14, leg. Messutat (cJM); 4 ex., Rhodes (FO 2), Kolymbia, beach, 26 Apr. [20]14, leg. Messutat (cJM); 1 ex., Rhodes (FO 3), Archipoli NE, 150 m, 26 Apr. [20]14, leg. Messutat (cJM); 1 ♀, Rhodes (FO6), Laerma N, 260 m, 28 Apr. [20]14, leg. Messutat (cJM); 5 ex., Rhodes (FO 9), Soroni-Fanes, 30 m, 1 May [20]14, leg. Messutat (NMBE); 6 ex., N Peloponnesus, Kato Loutro, W Xilokastro, collected at night on the beach, 2 Jun. 2012, leg. P. Bialooki (cPB); 2 ♂♂ Platanos, NE Parnon Mts, 18 May 2012, leg. P. Bialooki (cPB); 1 ♀, Gerakini, 4 Jun. 1992, leg. V. Svilha (cJS); 1 ♂, Pelop. [onnese], Mt Taygetos, Asafigion, 24 Jun. [1974], Horak & Svihla leg. (cJS); 4 ♀♀, Nomos, Lakonias, 5 km S of Monemvasia, 13–25 May [19]89, leg. Rieger (cCG, cJM); 1 ♂, Makedonia-S, 6 km NE of Lithochoro, Plaka-coast, 40°07′32″ N, 22°33′03″ E, 30 m, 11 May 2005, leg. C. Bayer (NMBE); 1 ♂, 1 ♀, Rhodes, coll. Stöcklein (NMB); 2 ♀♀, Thessalien, Volo, coll. P. Meyer (NMB); 2 ♂♂, Parnass (NMB); 1 ♀, Taygetos, coll. Apfelbeck (NMB); 3 ex., Parnass, Paganetti [19]03, coll. Stöcklein (NMB); 1 ♂, 1 ♀, Kreta, Paganetti, coll. Stöcklein (NMB); 1 ♂, 1 ♀, Kreta, Agios Nikolao, 24 Apr. 1971, leg. Wittmer (NMB); 12 ex., E Peloponnesus, pass W Astros, 18 May 2012, leg. P. Bialooki (cPB); 1 ♀, Attica, Nea Makri N, 1–14 Jun. 2015, leg. Messutat (cJM); 1 ♀, Euboia, Mt Ochi, 2.5 km NE of Myli, 965 m, 4 Jun. 2015, leg. Messutat (cJM); 4 ex., Rhodes, Kolymbia, beach, 3 Apr. 2015, leg. Messutat (cJM); 2 ex., [Peloponnese] Nomos Lakonias, Strand, 5 km N of Monemvasia, 15–25 May [19]89, leg. Rieger (cJM); 15 ex., Rhodes, 5 May 1996, Paradissi Mts, 100 m, Paradisi env., R. Borovec leg. (cRB, BMNH); 2 ♂♂, 2 ♀♀, 5 ex., Rhodes, 10 May 1996, Filerimos Mts, Lalyssos, 267 m, R. Borovec leg. (BMNH); 4 ex., Attiki, Varkiza, 8 May 1993, leg. E. Colonnelli (BMNH); 1 ex., Creta, nom. Iraklion, Matala, 13–15 Jun. 1994, leg. Regalin (BMNH); 2 ♂♂, 2 ♀♀, Attika, Sounio, 50 m, 17 May 2011, leg. Bahr, Bayer, Brunner and Winkelmann (cHW); 3 ex., Crete Chania, Potamida, 25 Apr. 2011, leg. H. Winkelmann (cHW); 1 ♀, Peloponnes, Kaliani, Prov. Korinthia, leg. W. Ziegler, 5 May 2010 (cHW); 1 ♂, Drama, Prosotsani, leg. W. Ziegler, 20 Jul. 2009 (cHW); 1 ♂, Korinthia, Stimfalia, 13 May 2007, leg. W. Ziegler (cHW); 1 ♂, Rhodos, Kallithea, 5–19 May 2005, leg. S. Scharf (cHW); 2 ♀♀, Kreta, Knossos Umgb., Jun. 1992, leg. H.-J. Grunwald (cHW); 3 ex., Platistomo, 31 May 2013, leg. Eifler (cHW); 1 ♂, Peloponnes, Argolis, Mt Didymo, Didyma doline, 6 Apr. 2010, leg. Winkelmann (cHW); 1 ♀, Peloponnese, Argolis, Palea Epidavros, 1–10 Apr. 2010, leg. Winkelmann (cHW); 1 ♀, Peloponnese, Corinthia, Korfos env., 10 Apr. 2010, leg. Winkelmann (cHW); 1 ♀, Peloponnes, Galatas, 7–28 Jun.[1976], leg. Köstlin (SDEI); 1 ex., Ost-Kreta, Vai, 6 Jun. 1999, leg. U. Heinig (SDEI); 1 ♀, Süd-Kreta, Umgb. Koutsouras, 1 Jun. 1999, leg. U. Heinig (SDEI); 1 ♂, Kreta, Umgb. Matala, 17–23 Apr. 2001, leg. Sieber (SDEI); 4 ex., Kreta, May [19]87, Chersonnissos, leg. R. Frieser (SDEI); 1 ex., Athen, [leg.] Schulz (SDEI); 1 ♂, Crete, Heraklion, Timbaki, 3 Apr. 2013, leg. H. Winkelmann (cHW); 1 ♀. Rhodes, Afantou env., 200 m, 10 May 1996, leg. R. Borovec (cRB); 2 ♀♀, 13 ex., Rhodos, Umgb. Faliraki, E [end of] May [19]83, R. Frieser leg. (SDEI); 1 ♂, Kreta, Aghios, Nikolaos 12 Apr.–3 May 1975, leg. R. Köstlin (SDEI); 1 ♀, Peloponnes, Astros, 14 Jun. [19]86, leg. H. Nebauer (SDEI); 1 ♂, Rhodes, 7 May 1995, Koutsoutis env., pr. Emponas, 400 m, leg. Ing. K. Schön (cKS); 12 ex., Crete Isl., Rethymno, Agia Gallini, 0 m, 16 May 2009, leg. C. Germann (cCG, NMBE, NMSO); 1 ♂, Crete Isl., Heraklio, N Demati (E Pirgos), N35°02′28″ N, E25°18′01″ E, 160 m, 19 May 2009, leg. C. Germann (cCG); 11 ex., Kalymnos Island, 1–6 Jun. 2012, leg. T. Kissling and M. Albrecht (cCG, NMBE, NMSO); 1 ♂, Peloponnese, Voidokilia-Bucht, Sommer [summer] 2005, leg. K. Germann (NMBE); 7 ex. Samos, 3 km NE of Pythagoreio, 20 m, 10 Jun. 2014, leg. Fouque René (cPB); 28 ex., Xanthi, Nestos-Delta, N of Chrysoupoli, Ruderalfläche, 41°00′46″ N, 24°42′39″ E, 40m, 29 May 2016, leg. C. Germann (cCG, NMSO).

ITALY (first records for Italy): 13 ex., Sicilia mer. [meridionale], Cassibile, 80 m, 12–13 May 1995, beating of Amygdalus , leg. R. Borovec (cCG, cRB); 4 ex., Calabrica, leg. Baudi (SDEI); 1 ♂, 1 ♀, Sicilia mer., Cassibille [sic!], 80 m, 12–13 May 1995, beating of Amygdalus , leg. M. Holecova (SDEI).

TURKEY: 1 ex., Asiatic Turkey, 1934, leg. Nihal Shevket (BMNH); 1 ex., Didyma, 21 May 1995, leg. Dr. A. Hamet (cRB); 1 ♂, Smyrna, coll. Apfelbeck (NMB); 8 ex., E Selendi, S Akhisar, 17 May 2010, leg. P. Bialooki (cCG, cPB); 5 ♂♂, 7 ♀♀, N Izmir, 8 km N of Bergama, 22 May 2011, leg. P. Bialooki (cPB, cCG); 2 ♀♀, S Antalya, Gündogmus, 27 Sep. 1980, leg. N. Lodos (cCG, cJF); 1 ♀, Menavgat, 5 May 1969, leg. Wewalka (cJF); 1 ♂, Smyrna [Izmir], coll. Apfelbeck (NMB).

Variability and differentiation

Males are more gracile than females. The colour of legs varies from dark brown and blackish to yellowish. The most similar species is Polydrusus calabricus which shares the body shape (long-oval elytra) and the shape of the penis ( Fig. 7F–G ). Both are easily separable by the shape of the eyes (see key and Fig. 3C–F ). Only P. rhodiacus shows similar eyes ( Fig. 2E–F ), but differs in legs deprived of scales and the elongate shape of the penis ( Fig. 7D ).

Ecology

Collected from April to June (exceptionally also in September) at various altitudes from beach to montane level from different deciduous trees ( Amygdalus , Pistacia lentiscus , Prunus , Quercus ) and Cupressaceae .

Figures

Fig. 3. A–B. Habitus of Polydrusus (Conocetus) crinipes sp. nov. A. Holotype, male from Corfu, Greece. B. Female from Bulgaria. — C–D. Polydrusus angustus (Lucas, 1854). C. Male from Rhodes, Greece.D. FemalefromKalymnos,Greece.— E–F.Polydrususcalabricus(Faust,1890).E.Malefrom Peloponnese, Greece.F. Female from Cephalonia, Greece.

Fig. 7. A–H. Penis of Polydrusus (Denticonocetus) and P. (Conocetus) spp., dorsal and lateral views. A. P. (D.) kahri Kirsch, 1865, Italy, Sicily. B. P. festae (F. Solari, 1925), Greece, Samos. C. P. stierlini (Schilsky, 1910), Greece, Macedonia.D. P. rhodiacus (Schilsky, 1912),Cyprus, Dionios. E. P. crinipes sp. nov., Greece, Corfu. F. P. angustus (Lucas, 1854), Greece, Crete. G. P. calabricus (Faust, 1890), Greece, Peloponnese. H. P. bardus Gyllenhal, 1834, Greece, Crete.

Fig. 11. Map showing records of five species of Polydrusus (Conocetus) and of P. (incertae sedis) longus (Stierlin, 1884) (map copyright: Google).

Fig.9.A–C.InternalsacofPolydrusus(Denticonocetus)kahriKirsch,1865.A.Italy,Sicilia.B.Croatia, Istria. C. Italy, Monte Baldo. D–P. Internal sac, dorsal and lateral views of: D. P. (Conocetus) festae (F. Solari, 1925), Greece, Samos. E. P. stierlini (Schilsky, 1910), Greece, Poligiros. F. P. rhodiacus (Schilsky, 1912), Cyprus, Dionios. G. P. crinipes sp. nov., Greece, Corfu. H. P. angustus (Lucas, 1854), Greece, Crete. I. P. calabricus (Faust, 1890), Greece, Patra. J. Ditto, Italy, Capo Leuca. K. P. bardus Gyllenhal, 1834, Greece, Crete. L. Ditto Iran, Lorestan. M. P. marcidus Kiesenwetter, 1864,Turkey,Antalya.N. P. baudii(Faust, 1889),Cyprus.O. P.transjordanussp. nov.,Jordania,Ajlun. P. P. grandiceps (Desbrochers des Loges, 1875), lectotype, Lebanon.

Fig.2.A–B.HabitusofPolydrusus(Conocetus)grandiceps(DesbrochersdesLoges,1875).A.Lectotype male from Lebanon. B. Female from Syria.— C–D. Polydrusus transjordanussp. nov., paratypes from Jordan. C. Male. D. Female. — E–F. Polydrusus rhodiacus (Schilsky, 1912). E. Male from Nicosia, Cyprus.F. Female lectotype from Cyprus.