Rattus sanila Flannery and White 1991

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Muridae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press, pp. 1189-1531 : 1487

Rattus sanila Flannery and White 1991

Rattus sanila Flannery and White 1991 , Nat. Geog. Res. Explor., 7: 102.

Type Locality: Bismarck Arch., New Ireland, Balof, site 2.

Vernacular Names: New Ireland Forest Rat.

Distribution: Apparently endemic to New Ireland.

Discussion: Rattus species group unresolved. Represented only by subfossil fragments dated at 3000 years before present and older (late Pleistocene to mid-Holocene), but may still occur in primary forest, which has not been adequately sampled. Flannery and White (1991) described sanila as a subspecies of R. mordax , but most dental measurements of sanila exceed and do not overlap those of even the largest known R. mordax , suggesting the former to be a separate species, which even Flannery and White acknowledged (also see Flannery, 1995 b). Recent taxonomic and stratigraphic analysis of rodent remains from New Ireland and Manus Isl has confirmed the specific status of sanila (K. P. Aplin and M. Leavesley, in litt., 2004). Although originally phylogenetically allied with R. mordax ( Flannery and White, 1991), K. Aplin (in litt., 2004) noted that upper molars of R. sanila "... are very complex in comparison with typical New Guinea Rattus ," and he suspects the species may be a relict of an archaic, and original, dispersal of ancestral Rattus stock to New Guinea and Australia.