Clistopyga marcapatensis Bordera & Palacio, 2019

Bordera, Santiago & Palacio, Edgard, 2019, The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part IV: the C. eldae species group, with the description of three new species, Zootaxa 4564 (2), pp. 327-346 : 337-338

Clistopyga marcapatensis Bordera & Palacio , sp. nov.

( Figs 1C , 2C , 3C , 4C , 7 A–G)

Diagnosis. Clistopyga marcapatensis sp. nov. may be distinguished from all other species of the C. eldae species group by the combination of the following characters: face with fine and moderately dense setiferous punctures ( Fig. 7C ); occipital carina only weakly insinuated dorsally or absent ( Figs 7A, B, D, E ); submetapleural carina strong and complete ( Fig. 4C ); hind wing with distal abscissa of Cu 1 weakly pigmented, joining cu -a + Cu 1 clearly below centre ( Fig. 3C ); tergite I 1.86–1.95× as long as posteriorly broad smooth and shiny, with scattered setiferous punctures ( Fig. 2C ); ovipositor weakly up-curved at distal 0.3–0.4 ( Fig. 7F ), about same length as hind tibia; ovipositor sheath about 0.78–0.8× as long as hind tibia; ventral two thirds of gena white ( Fig. 7A, E ).

Description. Female: Body length 6.2–6.6 mm (without ovipositor). Fore wing length 4.3–4.7 mm.

Head ( Figs 7 A–E). In dorsal view, moderately narrowed behind eyes. Gena smooth and shiny with very sparse setiferous punctures, in dorsal view rounded, 0.23–0.25× as long as eye ( Fig. 7B ), in frontal view straight and strongly constricted below eyes ( Fig. 7C ). Frons smooth and shiny. Vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 0.91–1.0× its maximum diameter. Distance between hind ocelli 0.82–0.91× the maximum diameter of posterior ocellus. Occipital carina weakly insinuated dorsally ( Figs 7A,B, D ) or absent. Face with fine and moderately dense setiferous punctures, distance between punctures more than three times the diameter of punctures. Clypeal suture moderately curved ( Fig. 7C ). Clypeus 1.71–1.80× as broad as medially long, weakly convex, dorsally, slightly concave ventrally, apical margin something bilobed. Malar space 0.43–0.44× as long as basal mandibular width, with a short granulate stripe between eye and mandible. Antenna with 29–30 flagellomeres, first flagellomere 7.60–7.83× as long as wide.

Mesosoma ( Figs 1C , 3C , 4C , 7 D–E). Pronotum smooth and shiny with fine sparse setiferous punctures on upper margin and hind corner. Epomia absent ( Fig. 7E ). Mesoscutum smooth and shiny, with very fine and sparse setiferous punctures. Notauli very weak, reaching about 0.4 of length of mesoscutum ( Fig. 7D ). Mesopleuron shiny, with very sparse setiferous punctures, except in posterior part under speculum. Epicnemial carina strong, its dorsal end straight, reaching a little below of centre of pronotum ( Fig. 7E ). Metapleuron shiny, glabrous, except for some sparse setiferous punctures at posterior part, 1.67–1.77× as long as deep. Submetapleural carina strong and complete ( Fig. 4C ), somewhat elevated at the anterior part. Propodeum smooth and shiny, with sparse and fine setiferous punctures laterally ( Fig. 4C ); in dorsal view 1.0–1.1× as long as medially wide. Propodeal spiracle at groove separating propodeum and metapleuron, groove interrupted by spiracle ( Fig. 4C ). Hind leg with femur 3.58–3.86× as long as deep, 0.85–0.90× as long as tibia. Fore wing with vein cu-a opposite Rs & M. Vein 2 rs-m about 0.36–0.47× as long as abscissa of M between 2 rs-m and 2 m-cu. Abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.33–1.42× as long as Cu 1 b. Hind wing with vein cu-a 0.25–0.39× as long as first abscissa of Cu 1. Vein cu-a reclivous, first abscissa of Cu 1 vertical ( Fig. 3C ). Distal abscissa of Cu 1 well pigmented.

Metasoma ( Figs 1C , 2C , 7F, G ). Tergite I 1.86–1.95× as long as posteriorly broad, smooth and shiny, with fine and sparse setiferous punctures laterally ( Fig. 2C ); spiracle near its basal 0.4; lateromedian longitudinal carinae strong, reaching about 0.25 of length of tergite; lateral longitudinal carinae strong, almost reaching spiracle. Sternite I extending back about 0.8 of length of tergite ( Fig. 7F ). Tergite II 1.08–1.19× as long as posteriorly broad, central region shiny, with fine and moderately dense punctures ( Fig. 2C ); rest of tergites shiny, progressively more densely and strongly punctate ( Fig 7G ). Ovipositor slender, matt, slightly upcurved at distal 0.4 ( Fig. 7F ), same length as hind tibia. Ovipositor sheath about 0.8× as long as hind tibia, length of setae on average 1.5–1.6× the sheath basal width.

Colouration ( Fig. 1C , 7 A–G). Head black with clypeus, face, frontal and vertical orbits, mouth parts except apex of mandibles, malar space and ventral two/third of gena, white; antenna with scape and pedicel dark brown dorsally, widely white at ventral side, flagellomeres brown, basal flagellomeres whitish on the ventral side. Mesosoma mostly yellow, darker dorsally; propleuron, tegula, subalar prominence, anterior part of pronotum, ventral part of mesopleuron and posterior part of metapleuron, whitish; scutellum and postscutellum yellow; posterior rim of propodeum black. Metasoma mostly dark brown, anterolateral marks of tergites II, anterior part of tergites III–VI extending back laterally, and posterior rim of tergites I–VIII, whitish. Legs predominantly white; fore leg with femur tibia and tarsi yellow; mid leg with base of trochanter, dorsal narrow stripe on femur and dorsal basal and distal part of tibia brown, tarsi infuscate; hind leg with anterior wide mark on coxa, anterior and posterior basal marks on trochanter, basal part, dorsal stripe and anterior longitudinal wide stripe on femur, tibia except mid part, and distal part of tarsi, brown. Wings hyaline, pterostigma dark brown. Ovipositor sheaths dark brown. Ovipositor reddish.

Male: unknown.

Etymology. The specific name refers to Marcapata district (Cusco, Perú), area where the type material was collected.

Type material. Holotype. Perú: 1 ♀, Quincemil , 750, nr. Marcapata, 20-30 Oct. 1962, Luis Peña, (AEIC).

Paratypes. Perú: 1 ♀, Avispas , 30 m, nr. Marcapata, Sept. 1962, Luis Peña ( AEIC) ; 1 ♀, same locality, 1-15 Oct. 1962, Luis Peña ( AEIC) .

Distribution. Peru.

Figures

FIGURE 1. Habitus of Clistopyga spp., females: A—C. eldae (paratype); B—C. catarina (holotype); C—C. marcapatensis (holotype); D—C. polita (holotype). Scale bar: 1 mm.

FIGURE 2. Morphology of Clistopyga spp., tergites I–III, females: A—C. eldae (paratype); B—C. catarina (paratype); C—C. marcapatensis (holotype); D—C. polita (holotype). Scale bar: 0.3 mm (A); 0.5 mm (B–D).

FIGURE 3. Morphology of Clistopyga spp., hind wing, females: A—C. eldae (paratype); B—C. catarina (paratype); C—C. marcapatensis (paratype); D—C. polita (holotype). Scale bar: 0.3 mm.

FIGURE 4. Morphology of Clistopyga spp., metapleuron, females: A—C. eldae (paratype); B—C. catarina (paratype); C—C. marcapatensis (holotype); D—C. polita (holotype) (arrow: submetapleural carina). Scale bar: 0.3 mm.

FIGURE 7. Morphology of Clistopyga marcapatensis, female: A–C, head; A—lateral view (holotype), B—dorsal view (paratype), C—frontal view (paratype), D–E, head and mesosoma; D—dorsal view (paratype), E—lateral view (holotype). F– G, metasoma (paratype); F—lateral view, G— dorsal view. Scale bar: 0.3 mm (A–D); 0.5 mm (E, F).