Incoltorrida benesculpta, Perkins & Bergsten, 2019

Perkins, Philip D. & Bergsten, Johannes, 2019, New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae), Zootaxa 4657 (1), pp. 57-96 : 72-73

Incoltorrida benesculpta , new species

Figs. 8 (habitus); 16, 20 (genitalia); 35 (map); 42, 43 (habitat)

Type Material. Holotype (male): Fianarantsoa, 3.2km S Ambohimanjaka , 20º 14.0343’ S 47º 5.59145’ E, waterfall and hygropetric habitat near hwy. 7, elev. 1415 m, 5 xi 2014, P. D. Perkins ( NHRS) . Paratypes (19): Same data as holotype (13: NHRS, MCZ, BMNH, PBZT / MBC); Antsiranana, Sava, Mad.; Antsirana ; Marojejy NP, 800m N from Camp I, Humbert waterfall; hygropetric; 14.4333S 49.773E; MAD 14-48 , ex. bedrock pools at side of waterfall, elev. 550 m, 8 xi 2014, P. D. Perkins ( MAD 14-48 ) (6 MCZ) .

Differential Diagnosis. Together with I. galoko the smallest known member of the genus (l/w ca. 1.85/1.05). The elytral costae are high and strong, and are more sinuate than in other species. The serial punctures are large and in sulcate grooves. The sculpture of the pronotum is similar to that of I. quintacostata , but I. benesculpta has a more distinct midlongitudinal carina in the basal half, and the sides of the pronotum in the anterior half are more arcuate. The elytra and the aedeagi of the two species are quite dissimilar.

Description. Size: holotype (length/width, mm): body (length to elytral apices) 1.85/1.05; head width 0.40; pronotum 0.40/0.81; elytra 1.17/1.05. Dorsum black, venter dark brown, legs brown except femoral-tibial articulations and tarsi black.

Sides of frontoclypeal shield slightly arcuate toward the midline, such that apical part is slightly wider than the width at midlength, and slightly narrower than the posterior area of the frons.

Short oblique carinae on anterior 1/3 of pronotum are well developed, no indication of midlongitudinal carina. The basal 1/3 has a short carina on each side, and a very low narrow midlongitudinal carina.

Elytron with eight strong high costae; fourth costa interrupted with strong punctures; fifth costa weakly bisinuate; sixth and seventh costae very indistinct. Serial punctures large and distinct.

Sides of metaventral tabella more distinctively arcuate than in other species. Midlongitudinal groove in the posterior 1/2 of metaventral tabella gradually widens from posterior to anterior.

Midlongitudinal carina of first abdominal ventrite strong and extends length of ventrite.

Male genitalia very short, with the apical part narrowed and spinose ( Figs. 16 , 20 ).

Etymology. Named in reference to the markedly sculptured pronotum and elytra.

Figures

FIGURE 8. Incoltorrida benesculpta, habitus views of holotype.

FIGURES 14–18. Incoltorrida spp.; (14) I. madagassica, male genitalia of holotype (after Steffan 1973, permanent slide mount). (15–17) male genitalia apex showing pores of I. madagassica and spines of I. benesculpta and I. quintacostata; (18) I. quintacostata, aedeagus of holotype.

FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype.