Troporhaconotus, Belokobylskij & Zaldívar-Riverón, 2021

Belokobylskij, Sergey A. & Zaldívar-Riverón, Alejandro, 2021, Reclassification of the doryctine tribe Rhaconotini (Hymenoptera, Braconidae), European Journal of Taxonomy 741, pp. 1-168 : 156-158

Genus Troporhaconotus gen. nov.

urn:lsid:zoobank.org:act:6D821B9C-95EC-4CAC-873F-1FA1F40902A6

Rhaconotus jacobsoni species group – Belokobylskij 2001: 134. — Qanh & Long 2019: 37.

Etymology

From “tropics”, the climatic region where this taxon appears to be restricted, and the generic name “ Rhaconotus ”. Gender: masculine.

Type species

Pseudospathius jacobsoni Szépligeti, 1908 .

Description ( Figs 58–59 )

HEAD. Head subglobular. Vertex densely and finely granulate or granulate-reticulate. Ocelli arranged often in almost equilateral triangle. Frons almost flat and without median keel. Eyes with distinct emargination opposite eyes, glabrous. Occipital carina complete dorsally, not joining ventrally with hypostomal carina being obliterated at short distance. Malar suture absent. Clypeal suture distinct and complete. Clypeus with more or less distinct ventral flange. Hypoclypeal depression medium size, round. Postgenal bridge rather wide. Palps rather short; maxillary palps 6-segmented, labial palps 4-segmented. Scapus wide and rather short, without apical lobe or basal constriction. First flagellar segment subcylindrical, almost straight, always longer than second segment.

MESOSOMA. Mesosoma long and narrow, 2.5–3.0 times as long as its high. Pronotum dorsally distinctly convex, with more or less distinct pronotal carina in anterior 0.3–0.4. Mesonotum not highly and weakly gently-roundly elevated above pronotum (lateral view); its all three lobes medially widely glabrous. Notauli complete, deep, crenulate. Prescutellar depression relatively long, with median carina, rugulose. Scuto-scutellar suture rather distinct. Scutellum with distinct lateral carinae at least in anterior half. Metanotum often with long and acuminate median tooth (lateral view). Sternaulus (precoxal furrow) distinct, relatively deep, long, running along all length of lower part of mesopleuron, distinctly crenulate. Prepectal carina complete, enlarged below opposite fore coxa. Postpectal carina absent. Propodeum at least with long median and two lateral carinae, often basolateral areas not delineated posteriorly, areola indistinct; lateral tubercles and propodeal bridge absent.

WINGS. Pterostigma bicolorated, mostly dark brown with pale its basal part. Radial vein (r) arising usually from or more or less behind middle of pterostigma. Radial (marginal) cell not shortened. Both radiomedial veins (2-SR, r-m) present. Recurrent vein (m-cu) distinctly postfurcal. Discoidal (discal) cell petiolate, narrow and long. Nervulus (cu-a) weakly postfurcal or sometimes subinterstitial. Mediocubital vein (M+CU1) rather distinctly curved distally to anal vein (2-1A). Parallel vein (CU1a) always interstitial. Brachial (subdiscal) cell closed usually before or (rarely) on level of recurrent vein (m-cu). Hind wing with three hamuli. Radial (marginal) cell without additional transverse vein (r). Medial (basal) cell rather narrow, but distinctly widened apically. Nervellus (cu-a) present. Submedial (subbasal) cell small. First abscissa of mediocubital vein (M+CU) distinctly shorter than second abscissa (1-M). Recurrent vein (m-cu) present, but often not pigmented.

LEGS. Fore tibiae with several distinct spines arranged in almost single line. Middle leg short and with short tarsal segments. Hind coxa with distinct basoventral corner and distinct, but small tubercle. Hind femur with rather distinct dorsal protuberances; femur of male distinctly broaded. Hind tibia with long, more or less dense and semi-erect setae. Hind basitarsus 0.6–0.7 times as long as second–fifth segments combined. Legs usually with dark and pale colour pattern.

METASOMA. Six dorsally visible and hardly sclerotised tergites. First tergite not petiolate, but very long, its dorsope rather large, basolateral lobes absent. Acrosternite of first segment 0.15–0.2 times as long as first tergite, its apical margin situated distinctly before level of spiracles. First and second tergites not fused immovably. Second tergite without basal area delineated posteriorly by furrow; apical area of second tergite present, long, coarsely sculptured, delineated anteriorly by relatively shallow and wide crenulate furrow and posteriorly by rather deep and wide curved second suture. Second–sixth tergites with separated laterotergites. Sixth tergite of female always enlarged, distinctly longer than fifth tergite, covered following apical segments, with usually distinctly convex its posterior margin, but only sometimes with more or less distinct incurvation posteriorly.

Diagnosis

This new genus differs from the remaining Rhaconotini genera by having more than five dorsally visible metasomal tergite and with distinctly enlarged sixth tergite covering following tergites, first metasomal tergite long, more than 2.0 times as long as its apical width, second tergite without basal area and always

with apical lenticular area rather distinctly delineated anteriorly by deep and rather wide furrow, and mesosoma long with mesoscutum weakly and gently elevated above pronotum.

Composition

Troporhaconotus anivoranensis ( Granger, 1949) comb. nov. (AF); T. asulcus (Shi & Chen, 2004) comb. nov. (OR); T. ceylonicus (Belokobylskij, 2001) comb. nov. (OR); T. chinensis ( Belokobylskij & Chen, 2004) comb. nov. (OR); T. directus (Long, 2019) comb. nov. (OR); T. glaphyrus ( Chen & Shi, 2004) comb. nov. (OR); T. jacobsoni ( Szépligeti, 1908) [= Rhaconotus filicornis ( Enderlein, 1912) ] comb. nov. (OR); T. laevigatus (Long, 2019) comb. nov. (OR); T. longithorax (Belokobylskij, 2001) comb. nov. (OR); T. robustus (Long, 2019) comb. nov. (OR); T. simulatus (Long, 2019) comb. nov. (OR); T. thayi Belokobylskij, 2001 comb. nov. (OR).

Hosts

Unknown.

Distribution

Afrotropical and Oriental regions.

Remarks

In Jasso-Martínez et al. (2019), the only included species of this genus, T. thayi (Belokobylskij, 2001) , was nested at the base of a large clade with taxa having mainly five-tergited metasoma. Despite its absence of sharp morphological diagnostic characters, the above combination of features together with its phylogenetic affinities allowed us to propose the erection of this new genus.

Figures

Fig. 58. Troporhaconotus jacobsoni (Szépligeti, 1908) gen. et comb. nov., ♀, holotype, HNHM. A. Habitus, lateral view. B. Head, front view. C. Head, dorsal view. D. Head, lateral view. E. Basal segments of antenna. F. Fore tibia. G. Hind coxa and femur, lateral view. H. Hind tibia, lateral view. I. Mesosoma, lateral view. J. Mesosoma, dorsal view. K. Propodeum and first metasomal tergite, dorsal view.

Fig. 59. Troporhaconotus jacobsoni (Szépligeti, 1908) gen. et comb. nov., ♀, holotype, HNMN. A. Fore wing. B. Hind wing. C. Metasoma, dorsal view. D. Metasoma, lateral view.