Exocelina atowaso (Shaverdo, Sagata & Balke, 2005)

Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael, 2012, Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 250, pp. 1-76 : 7

5. Exocelina atowaso (Shaverdo, Sagata & Balke, 2005) Fig. 25

Papuadytes atowaso Shaverdo, Sagata & Balke, 2005: 275.

Type locality.

Papua New Guinea: Madang Province, river below Bundi, approximately 05°44.85'S, 145°16.83'E.

Type material examined.

Holotype: male "PAPUA NEW GUINEA Madang Pr. below Bundi, 500 m, 26IX2002, M. Balke (PNG 23)", "267 DNA M Balke" [green label] (BMNH). Additional material. Madang: 5 males, 4 females "Papua New Guinea: Madang, below Bundi, 500 m, 26.IX.2002 Balke & Sagata (PNG 023)" (NHMW, ZSM). East Sepik/Enga: 1 male "DNA M. Balke 4918", "Papua New Guinea East Sepik, Lembena, 198m, 3.ix.2009, 04 46.974S 143 56.995E, Ibalim & Plus (PNG 243)" (ZSM). 1 ex. "Papua New Guinea East Sepik, Lembena, 198m, 3.ix.2009, 04 46.974S 143 56.995E, Ibalim & Plus (PNG 243)" (ZSM). 1 male "PNG 246" [hw] (ZSM). 1 male "Papua New Guinea East Sepik, Lembena, 335m, 10.ix.2009, 04 56.859S 143 59.375E, Ibalim & Plus (PNG 250)" (ZSM).

Diagnosis.

Beetle middle-sized, piceous, with paler head, shiny (Fig. 25); pronotum with distinct lateral bead; male antennomeres simple; male protarsomere 4 with middle-sized, slender, evidently curved anterolateral hook; median lobe with very strong submedian constriction and small subapical processes in ventral view and with almost rounded apex bearing setae in lateral view; paramere without notch on dorsal side, with relatively dense, long, thin setae.

Additions and corrections to the description.

For the complete description and illustrations see Shaverdo et al. (2005).

Size: Beetle middle-sized (TL-H 3.9-4.1 mm, TL 4.2-4.5 mm, MW 2.05-2.3 mm).

Male: Protarsomere 4 with middle-sized, slender, evidently curved anterolateral hook. Median lobe with very strong submedian constriction and small subapical processes in ventral view and with almost rounded apex bearing setae in lateral view. Paramere without notch on dorsal side, with relatively dense, long, thin setae. See Figs 9, 14a, b in Shaverdo et al. (2005).

Female: Without evident differences in external morphology from male, except for abdominal sternite 7 without striae.

Distribution and habitat.

Papua New Guinea. The species is known from Madang, East Sepik, and Enga Provinces (Fig. 50). For the habitat description see Shaverdo et al. (2005).