Megaselia kaleybarensis, Khameneh & Khaghaninia & Disney & Maleki-Ravasan, 2019

Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, Zootaxa 4711 (1), pp. 1-50 : 5

Megaselia kaleybarensis n. sp.

( Figs 39–54 )

Material examined. Holotype male, East Azerbaijan province, Arasbaran forests, Kaleybar region, 38°51.548’N 46°59.007’E, 1783 m, open forest, 1.vii.2017, S. Khaghaninia (17, UCMZ—13-91) . Paratypes: 46 males, locality data as the holotype (2 UCZM, 44 ICHMM) .

Description. Male. Whole fly as Fig. 39 . Frons as Fig. 40 , with microtrichia restricted to edges ( Fig. 41 ). Cheek with 4 bristles and jowl with 2 that are longer and more robust. Postpedicels, palps and proboscis as Fig. 42 , with postpedicels lacking SPS vesicles ( Fig. 43 ). Labella with numerous small bristles on ventral faces ( Fig. 44 ). Notopleuron with 3 bristles, with no cleft in front of these, and mesopleuron with hairs ( Fig. 45 ). Scutellum ( Fig. 46 ) with an anterior pair of hairs and a posterior pair of bristles. Abdomen as Fig. 47 , with hairs on segments 3–6 of venter 6. Tergites 2 to 4 as Fig. 48 . Hypopygium as Figs 49–51 . Legs yellowish brown with a darker tip to hind femur. Fore tarsus with posterodorsal hair palisade on segments 1–4 and 5 a little longer than 4 ( Fig. 52 ). Dorsal hair palisade of mid tibia extends about 0.9 times its length. Hairs below basal half of hind femur longer than those of anteroventral row of outer half ( Fig. 53 ), being 0.08 mm long. Hind tibia with 14–15 moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings ( Fig. 54 ) 1.70–1.75 mm long. Costal index 0.43–0.44. Costal ratios 3.7–3.8: 2.20: 1. Costal cilia (of section 3) 0.13 mm long. Hair at base of vein 3 only 0.03 mm long. With 4 axillary bristles, the outermost being 0.13 mm long. Sc not reaching R1. Haltere knob pale.

Recognition. In the key to the males of British species ( Disney 1989) it runs to triplet 104; but this is in part revised by Disney & Häggqvist (2018), in which it runs to M. subnitida (Lundbeck) . However, it differs in a number of details considered below. For the rest of the world the Nearctic species M. cribella Borgmeier (1964) is similar, but its longer costal index, position of the anterolateral frontal bristles and the tapered hypandrial lobe of the hypopygium exclude this species. With regard to the differences of our species from M. subnitida the dorsal hair palisade of mid tibia extends 0.76–0.90 times its length contrasting with 0.61–0.62 times in M. subnitida . The hairs below the basal half of the hind femur are longer ( Fig. 53 ) than in M. subnitida ( Fig. 56 ) and the longer hairs on the lateral fifths of abdominal tergite 2 are more numerous and extend almost to the lateral margins ( Fig. 48 ), but in M. subnitida they are fewer and end well before the lateral edges and the posterolateral hairs are longer ( Fig. 55 ). The hypopygia seem very similar.

Etymology. Named after the Kaleybar region.

Figures

FIGURES 39–48. Megaselia kaleybarensis n. sp. male. 39, whole fly; 40, frons; 41, part of frons enlarged; 42, postpedicels, palps and proboscis; 43, postpedicel; 44, ventral face of proboscis; 45, mesopleuron and notopleuron; 46, scutellum; 47, abdomen; 48, tergites 1 to 4. Scale bars 20 μm.

FIGURES 49–54. Megaselia kaleybarensis n. sp. male. 49, hypopygium left face; 50, epandrium left face; 51, hypopygium right face; 52, front tarsus; 53, hind femur; 54, wing. Scale bars 20 μm.

FIGURES 55–56. Megaselia subnitida male. 55, abdominal tergites 1 to 4; 56, hind femur. Scale bars 20 μm.