Psilota tristis Klocker, 1924

Young, Andrew D., Skevington, Jeffrey H. & Steenis, Wouter Van, 2020, Revision of the Psilota Meigen, 1822 flower flies (Diptera: Syrphidae) of Australia, Zootaxa 4737 (1), pp. 1-127 : 79-87

Psilota tristis Klocker, 1924

(Species plate: Fig. 35 ; Map: Fig. 64B )

Psilota tristis Klocker, 1924: 54 . Type locality: Queensland, Australia

Diagnosis: Face black, pale pollinose, with dark or pale pile. Frons and vertex black pilose. Thorax black. Thoracic pile variable in colour. Hind leg simple. Calypter variable in colour. Halter variable in colour. Abdomen black. Tergite pile variable in colour. Body length: 7.0– 14.1 mm.

Description: MALE: Head: Head black, face pale pollinose. Face dark or pale pilose, frons and vertex black pilose. Antenna orange to dark brown. Postpedicel variable in shape. Eye densely brown pilose. Thorax: Thorax black. Legs mostly orange to entirely dark brown. Hind femur simple. Hind tibia simple. Pleuron densely grey pollinose to bare and shining. Pile pleuron variable in colour. Pile on scutum and scutellum variable in colour, but always at least partly black on scutellum (male only). Wing colourless. Wing microtrichia variable. Alula completely microtrichose. Halter variable in colour. Calypter variable in colour. Abdomen: Abdomen black. Pilosity of tergites variable in colour. Sternites black, with variably coloured pilosity. Genitalia: variable (see below).

FEMALE: Identical except for normal sexual dimorphism.

Specimens examined: AUSTRALIA: Australian Capital Territory: Bendora , -35.221814 149.169528, 13.ii.1952, S.J. Paramonov, ANIC29034513 (1♀, ANIC) Black Mountain , -35.266303 149.095464, 24.x.1977; 13.iii.1961; 13.x.1965; 15.xii.1963; 21.ii.1966; 18.i.1966; 21.xii.1962; 26.xi.1959; 28.x.1966; 30.x.1962; 30.x.1964; 30.x.1969; xii.1987, Z. Liepa; I.F.B. Common; M. Irwin, Malaise trap; light trap, ANIC29034495 ; ANIC29034540 ; ANIC29034868 ; ANIC29034518 ; ANIC29034932 ; ANIC29034867 ; ANIC29034517 ; ANIC29034545 ; ANIC29034551 ; ANIC29034520-29034521 ; ANIC29034534 ; ANIC29034476 ; ANIC29035781 (7♂♂, 7♀♀, ANIC) ; Blundell’s [Cabin], -35.292886 149.141800, 25.xii.1937; 11.x.1950; 24.x.1950, M. Fuller; Paramonov, ANIC29034811 ; ANIC29034496 ; ANIC29034467 ; ANIC29034444 ; USNM _ ENT249831-32 (2♂♂, 5♀♀, ANIC, USNM) ; Canberra, -35.282000 149.128686, 1–10.xii.1953;, 25.xii.1985, F.M. Hull; D.C.F. Rentz, CNC _ Diptera 253774; ANIC29034493 (1♂, 1♀, CNC, ANIC) ; Cotter River , -35.531650 148.875125, 23.xi.1950; 29.xi.1962, Paramonov; Z. Liepa, CNC _ Diptera 253689; ANIC29034445 (2♀♀, ANIC) ; Gibraltar Peak, - 35.466661 148.933336, 24.xi.1948, Paramonov, ANIC29034516 (1♀, ANIC) ; Mt. Gingera , Bancroft , -35.576694 148.779703, in swamp on Epacris , 19.i.1963, Mackerras, ANIC29035758 (1♀, ANIC) ; Mt. Pemberton , 2km E. Gilmore , -35.416667 149.133333, dry Eucalyptus woodland, 18.x.2003; 1.xi.1947, D.J. Ferguson; S.J. Paramonov, ANIC29035754 ; ANIC29034832 ; ANIC29034878 (3♂♂, ANIC) ; New South Wales: 1 km E. Forbes Creek, SE, -35.429478 149.502800, 874m, 11.xi.2012, D.J. Ferguson, CNC _ Diptera 253796 (1♀, ANIC) ; 1 km North of Guth- rie’s Creek, Kosciuzko National Park , -36.413086 148.375475, 1722m, 1.ii.1974, K.R. Norris, ANIC29034875 (1♂, ANIC) ; 1.5 km West Northwest Lansdowne via Taree , -31.780136 152.51311, Cassina blossom, 12.ix.1985, G. Williams, AustralianMuseumK410249 (1♀, AM); 12 miles. N.W. Milton , -35.164567 150.298497, 487m, 5.xi.1968, D.H. Colless, ANIC29034527 (1♀, ANIC) ; 20 miles South of Milton , -35.319178 150.439158, 30.ix.1952, S.J. Paramonov, ANIC29034835 ; ANIC29034876 (2♂♂, ANIC) ; 24 km W South Grafton , -29.616667 152.733333, 1.xii.1990, G. and A. Daniels, Legs removed for DNA analysis; GDCBReg_19734 (1♀, GDCB) ; 25 km W of South Grafton , -29.616667 152.733333, 5.ix.1981, G. and A. Daniels, GGDCBReg_19696 (1♂, QM) ; 3 km North of Lansdowne , -31.758753 152.534258, Acradenia euodiiformis , 19.ix.1993, G. Williams, AustralianMuseumK409935 (1♀, AM) ; 4 miles North of Bateman’s Bay , -35.658456 150.177744, 14.x.1952; 12.x.1956, S.J. Paramonov; E.F. Riek, ANIC29034870 ; CNC _ Diptera 253873; CNC_ Diptera 253874; ANIC29034475 (3♂♂, 1♀, ANIC) ; 4–10 miles North of Bateman’s Bay , -35.658456 150.177744, 14.x.1959, S.J. Paramonov, ANIC29034871 (1♂, ANIC) ; 5 km North of Bilpin near Kurrajong , -33.462750 150.572836, 1979, N.W. Rodd, N.W. Rodd Collec- tion (Date is ambiguous reads ”6.3.79”), AustralianMuseumK408029 (1♂, AM); 5 mi’s. NE Nerriga , -35.086408 150.170681, 27.xi.1962, I.F.B Common & M.S. Upton, ANIC29034471 (1♀, ANIC) ; 6 km Northeast of Bilpin near Kurrajong , -33.462750 150.572836, 1979, N.W. Rodd, Date is ambiguous reads ”10.9.79”, AustralianMuseumK410272-410273, AustralianMuseumK408026; AustralianMuseumK410003 (3♂♂, 1♀, AM); Bald Rock Na- tional Park -28.860781 152.055686, on Leptospermum minutifolium blossom, 6-1.xii.1983; 7.xi.1984, D.K. Yeates, UQIC_ Reg 94612–94617; UQIC_ Reg 94629; UQIC_ Reg 94742 (3♂♂, 5♀♀, QM); Barrington Tops National Park, Polblue Swamp , -31.956389 151.435833, 18.i.2010, S.A. Marshall, debu00322900 (1♀, DEBU); Barrington Tops , -31.935611 151.569264, viii.1928, M. Fuller, ANIC29034844 (1♂, ANIC) ; Batesman Bay Area , Rt. 1 , -35.714256 150.179539, 13.xi.1988, F.C. Thompson, USNM _ ENT_00034117 (1♀, USNM); Bellmore Falls , -34.633956 150.555914, 1948, S.J. Paramonov, Date is ambiguous reads ”12.11.48”, ANIC29034842 (1♂, ANIC) ; Bendalong, -35.219983 150.485778, 6.x.1968, G. Daniels, GDCBReg_19653 (1♂, QM); Berowra , -33.621622 151.150317, ANIC29035012 (1♂, ANIC) ; Bilpin, Blue Mountains , -33.500259 150.532989, 26.ix.1978, N.W. Rodd, NAustra- lianMuseumK408015; AustralianMuseumK410152; AustralianMuseumK408021 (3♀♀, AM); Bimberi , -35.572761 148.708658, 25.xi.1934, Fuller, ANIC29034837 (1♂, ANIC) ; Blackheath, -33.635592 150.283517, 17.xi.1919, G.H. Hardy, ANIC29035019 (1♂, ANIC) ; 23.xi.1919, G.H. Hardy, ANIC29034548 (1♀, ANIC) ; Bolaro State For- est, -35.700000 150.033333, 22.i.2010, M. Batley, AustralianMuseumK407883 (1♂, AM); Broadwater Nat. Pk ., - 27.357453 151.09904, 26.viii.1984, D.K. Yeates, UQIC_ Reg 94766 (1♂, QM); Budawang NP, Western Distributor Road, Carter’s Creek , -35.516250 150.059500, 200m, 14.xii.2003, A. Zwick, ANIC29035753 (1♂, ANIC) ; Budth- ingaroo Creek Kanangra Boyd National Park , -33.988542 150.088364, 26.xii.1977, G. Daniels, GDCBReg_19700 (1♂, GDCB); Bullawa Ck., Mt. Kaputar Nat. Pk., -30.233333 150.1, 17.i.1994, D.K. Yeates, UQIC_ Reg 94680 (1♀, QM); Bungendore , -35.248714 149.450603, 27.x.1968, J.C. Cardale, ANIC29034515 (1♀, ANIC) ; Catherine Hill Bay , -33.148439 151.617817, 12.iv.1949, S.J. Paramonov, CNC _ Diptera 253699; CNC_ Diptera 253900 (2♂♂, ANIC); Clarence, Blue Mountains , -33.482308 150.224264, 1982, N.W. Rodd, Date is ambiguous reads ”8.11.82”, AustralianMuseumK410469; AustralianMuseumK408024; AustralianMuseumK410283 (3♀♀, AM); Clyde Moun- tains near Braidwood , -35.546072 149.950333, 26.x.1960; 26.x.1965, S.J. Paramonov, CNC _ Diptera 253713- 253725; ANIC29034862–29034863 ; ANIC29035007 ; USNM _ ENT249829 ; ANIC29034526 (16♂♂, 1♀, ANIC) ; Collaroy, Sydney, -33.926375 151.125936, 1921, Health Dept. (Date is ambiguous reads ”8.9.21”), ANIC29034858 (1♂, ANIC) ; Como, -34.004500 151.063789, 4.iv.1975, ANIC29034923 (1♀, ANIC) ; Concord near Sydney, - 33.926375 151.125936, 7.xii.1968, R. Coveney, AustralianMuseumK406924 (1♂, AM); Conimbla National Park , -33.80547 148.48256, 7.x.2012, M. Batley, AustralianMuseumK410608 (1♀, AM); Coocumbac Island Nature Re- serve, Taree , -31.920678 152.454450, Floodplain forest landing near small pool of water on fallen log, 18.i.1994, G.&T. Williams,AustralianMuseumK410628 (1♀, AM); Deep Ck., Narrabeen , -33.719681 151.297125, 28.xii.1961, D.H. Colless, ANIC29034514 (1♀, ANIC) ; Duffy’s Forest , near Sydney , -33.926375 151.125936, 14.xi.1970, D.K. McAlpine, AustralianMuseumK410117; AustralianMuseumK410132; AustralianMuseumK410140; AustralianMuseumK410254; AustralianMuseumK410294 (1♂, 4♀♀, AM); Durras Bay , -35.645003 150.291439, 12.xi.1953; 29.ix.1959, Z. Liepa; F.M. Hull, CNC _ Diptera 255056-255057, ANIC29034869 (1♂, 2♀♀, ANIC, CNC) ; East Moree , -29.461931 149.852467, 14.xii.1961, E. Cheah, CNC _ Diptera 245382 (1♀,); East of Taree, Manning Point , -31.894431 152.658919, littoral rainforest on Acmena smithii blossom T.I., 8.xi.1990, G. Williams, AustralianMuseumK410238 (1♂, AM); Gibraltar Range National Park, Waratah track, -29.5 152.333333, on Leptospermum sp. blossom, 10–12.xi.1984, D.K. Yeates, UQIC_ Reg 94568 (1♀, QM); Grose Vale , -33.584000 150.674000, 1978, N.W. Rodd, AustralianMuseumK410271 (1♂, AM); Gundamaian National Park , -30.511594 152.995342, viii.1962, U.Q.I.C. Loan #1036, JSM5623, UQIC_ Reg 94562 (1♂, QM); Heathcote , -34.083333 151.016667, 31.x.1948, Crawford,L.D., VAIC70225 (1♂, VAIC) ; Heathcote, near Sydney, -33.926375 151.125936, 18.xi.1970, D.K. McAl- pine, AustralianMuseumK410141 (1♂, AM); Hornsby , -33.704900 151.099011, 20.xi.1932, Darlington, Australia, CNC _ Diptera 253779 (1♀, CNC); Jervis Bay , -35.035431 150.668869, 1.x.1952; 18.i.1949; 18.ix.1949; 2.x.1952; 18.ix.1951; 24.i.1963; 4.ix.1951, Paramonov; T.G. Campbell; I. Liepa; Z. Liepa, ANIC29034450 ; CNC _ Diptera 253870; ANIC29034505 ; USNM _ ENT249830 ; ANIC29035003 ; ANIC29034877 ; USNM _ ENT249826 ; CNC _ Diptera 253888 (5♂♂, 2♀♀, ANIC, USNM); Katoomba , -33.713131 150.313267, 10.x.1953; 14.ii.1952; 4.xii.1952; 20.i.1921; 31.xii.1956, F.M. Hull; G.H. Hardy, CNC _ Diptera 253773; AustralianMuseumK410587; Jeff _ Skevington _ Specimen 30409-30410; AustralianMuseumK407101; CNC9032 (3♂♂, 3♀♀, CNC, AM, MCZ) ; Killara, - 33.768642 151.163478, 1.ix.1935, M.F. Day, ANIC29034873 (1♂, ANIC) ; Kosciusko, -36.455897 148.263600, F.H. Taylor, ANIC29034841 (1♂, ANIC) ; Kosciuszko National park , 3.2km WSW of Thredbo, near Dead Horse Gap , -36.520833 148.268333, 1496m, over narrow stream in flowering grassland, 1–11.i.2002, C.L. Lambkin, N.T. Starick, ANIC29007126–27 (2♀♀, ANIC) ; Ku-ring-gai Chase Nat. Park , -33.637567 151.233900, 2.x.1972, A. and G. Daniels, GDCBReg_19726 (1♂, GDCB); Loftus, near Sydney , -33.926375 151.125936, 7.xi.1970, D.K. McAl- pine, AustralianMuseumK410126; AustralianMuseumK410143; AustralianMuseumK410165 (3♀♀, AM); Lorien Wildlife Refugee , 3 km North of Lansdowne, -31.758753 152.534258, Acradenia euodiiformis blossoms, 25.ix.1991, G. Williams, AustralianMuseumK407176 (1♀, AM); Mellong Ck , 12.8km S of Putty , -33.050000 150.683333, 23.xi.1978, G. Daniels, Daniels Collection, GDCBReg_19660 (1♀, GDCB); Mooney Mooney Creek, near Gosford , -33.437894 151.246467, 3.ix.1984, D.K. McAlpine, B.J. Day, AustralianMuseumK405939 (1♂, AM); Mount Ka- putar National Park , near Dawsons Springs , -30.284722 150.167500, 1.i.2010, S.A. Marshall, debu00318889 (1♀, DEBU); Mount Keira, near Wollongong Uni Obser , -34.402692 150.856469, 25.ix.1992, M. Gregory, White label ”30”, AustralianMuseumK408012 (1♂, AM); Mount Royal near Singleton , -32.174206 151.326167, 12.xi.1983, B.J. Day, K.C. Khoo, AustralianMuseumK410616 (1♂, AM); 13.xi.1983, B.J. Day, AustralianMuseumK408018 (1♀, AM); Mount Tomah, Blue Mountains , -33.549614 150.416033, 19.x.1988, N.W. Rodd, N.W. Rodd Collection, AustralianMuseumK410281(1♀, AM); Mount Victoria , -33.583333 150.250000, 29.ix.1934, D.F. W., ANIC29034859 (1♂, ANIC) ; Mt. Kaputar N.P., Bullawa Ck., -30.14 150.06, 16–19.i.1994, M.E. Irwin, D.K. Yeates, malaise trap, UQIC_ Reg 94676–94678 (2♂♂, 1♀, QM); Mt. Wilson , -33.506347 150.379069, 27.ii.1936, D.F. Waterhouse, ANIC29034523 (1♂, ANIC) ; N.E.[New England] National Park, near Entrance, -30.476022 152.660181, 29.xi.1960, C.W. Frazier, ANIC29035006 (1♂, ANIC) ; Nadgee NR. near Eden, -37.399378 149.908592, wet sclerophyll forest, 14–15.ii.1986, D.J. Bickel, AustralianMuseumK410253 (1♀, AM); Narrabeen , -33.719681 151.297125, 15.ix.1934, D.F.W., CNC _ Diptera 253700–253701 (2♂♂, ANIC); Norfolk Island , -29.040836 167.954711, 30.xi.1971, D. Kru- ger, ANIC29034482 (1♀, ANIC) ; Northbridge, Sydney, -33.926375 151.125936, 23.viii.1932, Fuller, ANIC29034451 (1♀, ANIC) ; Nullo Mountains , 20 miles NE Rylstone , -32.737947 150.226400, 11.xi.1950, T.G. Campbell; Para- monov, ANIC29034506 ; ANIC29034469 ; ANIC29034497 (3♀♀, ANIC) ; O’Hares Creek, Darkes Forest , -34.229969 150.908403, 2.ix.1992; 23.ix.1992, M. GregoryAustralianMuseumK408017, AustralianMuseumK408010; AustralianMuseumK408016 (2♂♂, 1♀, AM); Otford , -34.216667 151.008333, on Acmena smithi flowers, 25.xi.1995, A. Sundholm, A. Scott, ANIC29035778 (1♂, ANIC) ; Peats ridge, -33.333333 151.233333, on flowers, Leptospermum , 23.ix.1995, A. Sundholm, ANIC29035759 (1♀, ANIC) ; Petersham, -33.896100 151.154300, x.1927, M. Fuller, ANIC29035016 (1♂, ANIC) ; Port Macquarie , -31.433356 152.899886, 26.ix.1981, D.K. McAlpine & B.J. Day, AustralianMuseumK410124; AustralianMuseumK410618 (1♂, 1♀, AM); Roseville, Sydney , -33.926375 151.125936, 1915, Date is ambiguous reads ”3.1.15”, ANIC29035023 (1♂, ANIC) ; Royal National Park , -34.114342 151.046586, 28.xii.1970, D.K. McAlpine, AustralianMuseumK410620 (1♂, AM); on leaves of Xanthorrhoea , 4.xii.1970, D.K. McAlpine, AustralianMuseumK410256 (1♀, AM); Ruby Creek, near Stanthorpe , -28.608586 152.010250, 4–7.xi.1969, E.C. Dahms, Possibly in Queensland, QM _ REG._NO._T223217 (1♂, QM); Rutherford Ck., Brown Mtn, nr. Nimmitabal , -36.600000 149.416672, 24.xii.1974, G. Daniels, GDCBReg_19643 (1♂, QM); SYDNEY , -33.926375 151.125936, 10.x.1948, Crawford,L.D., Collected Sydney University campus, VAIC70224 (1♂, VAIC) ; Siding Spring Observatory, -31.272222 149.061111, summit, 17.x.1997, C. Lambkin, J. Skevington & S. Winterton, UQIC_ Reg 94749–94751 (3♂♂, QM); South Black Range , Tallaganda Nat. Pk., 15km from Hoskin- stown , -35.413333 149.535000, flowering heath, 21.ii.2004, C. Lambkin, N. Starick, ANIC29035777 (1♂, ANIC) ; South Black Range, east of Hoskinstown , -35.411111 149.534444, 1130m, 31.xii.2010, D.J. Ferguson, CNC _ Diptera 253795 (1♀, ANIC); Summit, Mount Gower, Lorde Howe Island , -31.588175 159.073681, 19.ii.1971, D.K. McAlpine, AustralianMuseumK407879 (1♀, AM); Sydney, Terry Hills , -33.926375 151.125936, 14.ix.1963, D.H. Colless, ANIC29034872 (1♂, ANIC) ; Sydney, -33.926375 151.125936, 16.x.1918; 24.xi.1918; 24.ix.1922; 8.ix.1921, G.H. Hardy; Health Dept; Deane & G. Mocatta, ANIC29034541–42 ; ANIC29034857 ; ANIC29034549 ; Jeff _ Skevington _ Specimen 30406; UQIC_ Reg 94597 (3♂♂, 3♀♀, ANIC, QM); Tibooburra, Cobham Lake , - 30.146856 142.087394, 17.xi.1949, S.J.K. Paramonov, ANIC29034937 (1♂, ANIC) ; Toronto, -33.016194 151.582583, ANIC29034504 ; ANIC29035020 (1♂, 1♀, ANIC) ; Wambelong Ck, Warrumbungle NP, -31.323000 149.027000, Malaise across creek bed, 21.i–9.ii.2009, S.L. Winterton, Malaise trap, CNC661779–661782 ; CNC669607 (2♂♂, 3♀♀, CSCA) ; Warrumbungle National Park , Fan Summit, -31.13 149.01 ; -31.130000 149.010000, hilltop, 27.iii.1999, S.A. Marshall, CNC _ Diptera 254194–254197 (4♂, DEBU); Warrumbungle Na- tional Park , -31.2743 148.989427, 17.x.1997, J. Skevington, S. Winterton, C. Lambkin, malaise trap, UQIC_ Reg 94755 (1♀, QM); Waterfall , -34.141986 150.992144, 30.xii.1933, Fuller, CNC _ Diptera 253703 (1♀, ANIC); Winburndale Nat. Reserve, NE. of Bathurst , -33.333164 149.789450, 6.xii.1974, J. Cardale, ANIC29034501 (1♀, ANIC) ; Woy Woy, -33.489839 151.319942, 8.ix.1924, ANIC29034533 (1♀, ANIC) ; [Royal] National Park, Gunda- maia, -34.083944 151.103617, 23.viii.1925, Mackerras, ANIC29034535 ; ANIC29034536 (2♀, ANIC) ; [Royal] Na- tional Park, Sydney, -33.926375 151.125936, 4.x.1925; 13.x.1948; 18.xi.1955, Paramonov; D.K. McAlpine, ANIC29034547 ; ANIC29034443 ; CNC _ Diptera 253884; CNC_ Diptera 253889–253891; CNC_ Diptera 253892; AustralianMuseumK410623 (5♂♂, 3♀♀, AM, ANIC); approximately 24 km North Northwest Gloucester , - 31.795028 151.949758, Acmena smithii rainforest, 5.xii.1990, G. & T. Williams, T.2., AustralianMuseumK410246 (1♀, AM); near Braidwood , -35.434689 149.802942, 31.x.1951, S.J. Paramonov, ANIC29034880 (1♂, ANIC) ; ANIC29034550 (1♂, ANIC) ; near Conspicuous Cliff, South of Nornalup , -35.004314 116.880339, fw. cr. behind dunes, 7.xi.1991, D. Bickel, AustralianMuseumK407910 (1♀, AM) ; Northern Territory: 17 km North of Pine Creek , -13.666058 131.836867, on Eucalyptus foelcheana , 22.xi.1974, E.M. Exley & R.I. Storey, UQIC_ Reg 94759 (1♀, QM); Bill’s Point, Mary River , -12.932286 131.809983, 3.viii.1929, I.M. Mackerras, T.G. Campbell, ANIC29034904 (1♂, ANIC) ; Gregory Nat’l Park, Humbert Track 1.1km S Bullita Camp Ground , -16.213056 130.419722, nr weeping Boab tree, 5–12.vi.2001, M.E. Irwin, F.D. Parker, C. Lambkin, CNC _ Diptera 255042; ANIC29035774 (2♀♀, ANIC) ; Queensland: 10 km N of Dunwich, Stradbroke Island , -27.412264 153.440086, 19.ix.1981, G. and A. Daniels, GDCBReg_19749 (1♀, GDCB); 14 km W by N of Hope Vale Mission , -15.294200 145.108236, 10.x.1980, D.H. Colless, ANIC29034449 (1♀, ANIC) ; 15km N. Dalveen, -28.351623 151.972712, on Angophora costata , 13.i.1981, J. & C.R. King, QM _ REG._NO._T221284 (1♀, QM) ; 18 km South of Ravenshoe , -17.771204 145.485042, 9.ix.1983, N.W. Rodd, AustralianMuseumK410131 (1♂, AM) ; 2 km W of Rainbow Beach , -25.966667 153.150000, rainforest, 24.ix.1994, G. & A. Daniels & C.J. Burwell, GDCBReg_19576; GDCBReg_ 19578–19579; GDCBReg_19581; GDCBReg_19584 (2♂♂, 3♀♀, GDCB); Acacia Ridge 3 mi. R.R., Brisbane , - 27.526975 153.040322, 26.ix–3.x.1953; 12.ix.1967, F.M. Hull; E.C. Dahms, CNC _ Diptera 253764; CNC_ Diptera 253760; QM_ REG._NO._T223176 (2♂♂, 1♀, QM, CNC); Amiens State Forest, nr. Stanthorpe , -28.606395 151.799206, 24–26.xi.1981, G. Daniels & M.A. Schneider, UQIC_ Reg 94895 (1♂, QM); Binna Burra, Lamington National Park , -28.195125 153.188792, 700m, 29.x.1986, I.F.B. Common, ANIC29035782 (1♂, ANIC) ; Blue Mtn., 4 km NE Stanthorpe , SE Qld. Summit , -28.633343 151.966683, 977m, 28.ix.1987, D.K. Yeates, UQIC_ Reg 94627 (1♂, QM); Bluff Range, Biggenden , -25.600000 152.050000, 7–10.x.1972; 18.viii.1972, H. Frauca, ANIC29034555 ; ANIC29034997 (1♂, 1♀, ANIC) ; Bribie Island , -26.9861 153.132525, 26.ix.1984; 29.viii.1918, G. Daniels; H. Hacker, UQIC_ Reg 94892; QM _ REG._NO._T223187 (2♂♂, QM); Brisbane Forest Pk ., -27.416667 152.833333, 9.ix.1995, M.A. Schneider, UQIC_ Reg 94772 (1♂, QM); Brisbane, Mount Coot-tha , -27.483333 152.950000, 170m, hilltop, 19.ix.2014; 9.ix.1972; 10.viii.1997, J.H., A.M. and A.W. Skevington; A.M. Hemmingsen; J. & A. Skevington & S. Winterton, hand collected, CNC451769 ; CNC451770 ; UQIC_ Reg 94775; (3♂♂, 1♀, QM, CNC); Bris- bane , -27.526975 153.040322, 15.ix.1913; 1.i.1927; 11.ix.1965; 18.ix.1965; 21.ix.1963; 18.ix.1914; 20.ix.1916; 20.viii.1918; 2.ix.1927; 22.xii.1953; 25.ix.1925; 29.viii.1916; 7.ix.1927, J. Mann; C.F. Ashby; S.J. Paramonov; F.H. Roberts; H. Hacker; G.H. Hardy; L. Pottenger, UQIC_ Reg 94815–94818; UQIC_ Reg 94820; ANIC29034553 ; ANIC29034855 ; ANIC29034828 ; ANIC29034530 ; ANIC29034538–29034839 ; QM _REG._NO._T223167; QM_ REG._NO._T223170–223172; QM_REG._NO._T223178–223183; QM_ REG._NO._T223190; ANIC29034554 ; AustralianMuseumK407687; AustralianMuseumK407611; AustralianMuseumK407221–407222; AustralianMuseumK407226–407229; AustralianMuseumK407233–407236; AustralianMuseumK407239; AustralianMuseumK407566–407568; AustralianMuseumK407570–407572; AustralianMuseumK407574–407578; Australian- MuseumK407583–407592–407594; AustralianMuseumK407596–407599; AustralianMuseumK407601; AustralianMuseumK407603–407604; AustralianMuseumK407606; AustralianMuseumK407612; AustralianMuseumK407685; AustralianMuseumK407693; AustralianMuseumK407696; AustralianMuseumK407699–407702; AustralianMuseumK407705; AustralianMuseumK407709–407711; AustralianMuseumK407714–407716; UQIC_ Reg 94809; JeffSkevingtonSpecimen#31370–31371; Jeff _ Skevington _ Specimen 30416 (42♂♂, 45♀♀, AM, ANIC, MCZ, QM); Brown Lake , N. Stradbroke Is ., -41.345000 147.352778, 21–24.ix.1984, D. Yeates, QM _ REG._NO._ T221276 (1♀, QM); Bunya Mountains , -26.884475 151.597200; -26.886078 151.596803, 1925, H. Hacker, Date is ambiguous reads ”10.12.25”; QM _REG._NO._T223511; QM_ REG._NO._T223169 (2♀, QM); Caboolture , - 27.066672 152.966667, 25.viii.1959, F.A. Perkins, QM _ REG._NO._T222171 (1♀, QM); Cairns District , -16.918678 145.777725, Dodd, ANIC29034491 (1♀, ANIC) ; Caloudra, -26.799000 153.133000, 17.viii.1934; 1.ix.1940; 23.viii.1933; 5.viii.1932, F.A. Perkins; C.F. Ashby; J.K. Guyomar, QM _ REG._NO._T221280; ANIC29034529 ; CNC _ Diptera 253775; UQIC_ Reg 94715; ANIC29034850 (4♂♂, 1♀, ANIC, CNC, QM) ; Camira, nr. Ipswich, - 27.634903 152.915636, 12.ix.1981, G. Daniels, GDCBReg_19733 (1♂, QM); Camp Milo, Cooloola , -26.043286 153.033894, 15–18.x.1978, G.B. Monteith, QM _ REG._NO._T223177 (1♀, QM); Carnarvon National Park, Mount Moffatt Summit , -25.059722 148.043889, 1097m, 29.xi.1997, J. Skevington, C. Lambkin, QM _ REG._NO._ T221267 (1♀, QM); Carnarvon National Park, Sugarloaf Mountain Summit , -24.898611 147.778611, 1.xii.1997, J. Skevington, C. Lambkin, QM _ REG._NO._T221279 (1♂, QM); Carnarvon National Park, “Fly Hill” summit near West Branch Camp , -24.971167 147.989000, 900m, 12.x.2002, J. Skevington, CNC _ Diptera 294990 (1♂, CNC); Cleveland , -27.533333 153.266667, viii.1924, G. Daniels, AustralianMuseumK407602 (1♀, AM); G.H. Hardy, AustralianMuseumK407708 (1♀, AM); Cooloola , -26.043286 153.033894, 17–28.viii.1912, E.C. Dahms, light trap, open forest, QM _ REG._NO._T223185 (1♂, QM); Culgoa Floodplain NP, Reedy Swamp , 7km NW Toulby Gate -28.915722 146.976925, 138m, 18.xi.2009, C. Lambkin, N. Starick, hand net, CNC _ Diptera 255048; QM_ REG._NO._T222151 (2♀♀, QM); Currumbin , -28.136000 153.483000, 8.iv.1954, K.R. Norris, ANIC29034490 (1♀, ANIC) ; D’Aguilar National Park, Mount Glorious, Greene’s Creek , -27.323522 152.762033, 9.xi.2014, J.H., A.M. and A.W. Skevington, CNC385040–385041 (2♀, CNC) ; D’Aguilar National Park, Mount Glorious , -27.327778 152.753611, 687m, rainforest, 15–20.ix.2014, Malaise trap, CNC457517 (1♀, QM) ; Dayboro, 0.5 km South , - 27.200000 152.816667, 18–19.ix.1999, S.G. Evans, QM _REG._NO._T223220; QM_ REG._NO._T223231–223232 (1♂, 2♀♀, QM); Dunalley , 16.x.1918, G.H. Hardy, ANIC29034852 (1♂, ANIC) ; Dunk Island , -17.944558 146.153289, viii.1977, G.H. Hardy, AustralianMuseumK407595 (1♂, AM); Durham Downs- Nokandra , -26.116933 149.076992, 13.xi.1949, S.J. Paramonov, ANIC29034938 (1♀, ANIC) ; Eungella National Park , Chelmer’s Road , - 21.132822 148.492683, 19.xii.2014, J.H., A.M. and A.W. Skevington, CNC451694 ; (1♂, CNC) ; Goondiwindi, - 28.539150 150.298089, ix.1933; ix.1935; ix.1953; ix.1965, G.H. Hardy, AustralianMuseumK407607; Australian- MuseumK407694;AustralianMuseumK407610;AustralianMuseumK407712;AustralianMuseumK407717–407719 (6♂♂, 2♀♀, AM); Illaweena St , Drewvale , -27.643333 153.063333, 40m, scribbly gum/heath, 9.ix.2003, QM _ REG._NO._T222137–222138 (1♂, 1♀, QM); Kuranda , -16.816878 145.633497, F.P. Dodd, ANIC29034981 (1♂, ANIC) ; Lake Broadwater 25 km SW Dalby , -27.350439 151.096633, on Leptospermum flavescens blossom, 18.x.1985, D. Yeates, UQIC_ Reg 94588; QM _ REG._NO._T222153 (2♂♂, QM); Lake Broadwater, nr Dalby, Qld site A , -27.350000 151.100000, 28.ix.1986, G. and A. Daniels, GDCBReg_19516 (1♂, QM); Lamington National park , -28.210000 153.127000, iii.1921; 23.i–3.ii.2009, G.H. Hardy; G. Monteith & F. Turco, QM _REG._NO._ T222139; QM_ REG._NO._T223164–223165 (3♀♀, QM); Lookout, Clark Ra., Cathu State Forest , -20.822833 148.541667, 820m, 17.i.1995, G. and A. Daniels, GDCBReg_13079; GDCBReg_19572–19573 (3♂, QM); Ma- hogany Forest, Mount Moffat National Park , -24.933333 148.066667, 1200m, 25.ii.1996, C.J. Burwell, S. Evans, light trap, QM _ REG._NO._T222136 (1♀, QM); Maleny , -26.762631 152.852242, 1965, C.F. Ashby, ANIC29034861 (1♂, ANIC) ; McIlwraith Ra. 8km NE Coen , -13.888333 143.253500, 540m, rain forest, 7.i.1995, G. and A. Daniels, GDCBReg_13077–13078 (2♀♀, GDCB); Mossman, N.Q ., -16.450000 145.373333, Raqment, Name and date un- clear, CNC _ Diptera 253777 (1♀, CNC); Mount Elna, Summit , -23.159722 150.451667, 284m, 19.ix.1999, J.&A. Skevington, QM _ REG._NO._T221278 (1♂, QM); Mount Greville near Aratula , -28.083333 152.500000, 5.xii.1996, C.J. Burwell, S. Evans, QM _ REG._NO._T223210 (1♂, QM); Mount Marlay, nr. Stanthorpe , -28.655786 151.947126, on Leptospermum ; 22.ix.1985; 7.x.2002, D.K. Yeates, UQIC Loan UQIC_ Reg 94669; UQIC_ Reg 94736–94738; UQIC_ Reg 94732; ANIC29035755–57 ; UQIC_ Reg 94579; UQIC_ Reg 94671; CNC _ Diptera 253688 (10♂♂, 1♀, QM); Mount Tinbeerwah, nr Cooroy , -29.950000 152.433333, 29.ix.1985, G. and A. Daniels, GDCBReg_19559– 19560 (2♀♀, GDCB); Mount Walsh National Park, Bluff Range Biggenden , -25.567886 152.048761, 762m, 12.viii.1972; 4.x.1976, H. Frauca, CNC _ Diptera 253885, ANIC29034998 ; ANIC29035000 ; ANIC29035004 (4♂♂, ANIC) ; Mount Walsh summit , -25.583333 152.050000, 640m, open forest & rock slabs, 2–3.xii.1998, C.J. Burwell, S.G. Evans, QM _ REG._NO._T223218 (1♂, QM); Mt Macartney, Cathu State Forest , -20.834333 148.537333, 850m, rain forest, 17.i.1995, G. and A. Daniels, GDCBReg_13076 (1♀, GDCB); Mt. Coot-tha Reserve, Summit Lookout , -27.4847629 152.9591569, 19.ix.2014, J.H., A.W., & A.M. Skevington, CNC372634–372635 (2♂♂, CNC) ; Mt. Glorious , -27.333422 152.766644, 19.xii.1962; 25.iv.1930, C.F. Ashby, ANIC29034473–74 ; CNC _ Diptera 253864; QM_ REG._NO._T221270 (4♀♀, ANIC, QM); Mt. Walsh Nat. Park, Mt. Walsh summit , -25.573333 152.051389, 2–3.xii.1998, C. Lambkin, D. White, D. Yeates, S. Evans, C. Burwell, QM _ REG._NO._T221262– 221264(3♂♂, QM); Mt.Walsh,nr. Biggenden , -25.567886 152.048761, 1000m, 8.x.1976,H. Frauca, ANIC29034499 ; ANIC29034507 ; USNM _ ENT_00249827 (2♂♂, 1♀, ANIC; USNM); Noosa , -26.364544 152.967669, 1969, J.K. Guyomar, Date is ambiguous reads ”4.10.69”, ANIC29034864 (1♂, ANIC) ; Northeast of Coongara Rock , Ban Ban Range, Southwest Biggenden , -25.512597 152.037756, 24.viii.1976, H. Frauca, ANIC29035013 (1♂, ANIC) ; Nr. Rosedale, -24.631128 151.913664, 27.x.1974, H. Frauca, ANIC29034441 ; ANIC29034508 (2♀, ANIC) ; on blos- soming dogwood, Jacksonia scoparia , 27.x.1974, ANIC29034466 (1♀, ANIC) ; Oakwells site 7, -25.806667 148.271667, 580m, stock through, partly cleared gully, brigalow, vine scrub, 1.x.2005, C. Eddie, (1♂, QM); Orien- tos—Nappamerrie , -27.600878 141.106858, 5.xi.1949, S.J. Paramonov, ANIC20934902 ; CNC _ Diptera 255081 (2♂♂, ANIC); Rockpool Gorge, Mt. Walsh National Park, Biggenden , -25.605889 152.027519, 4.x.1976, H. Fr- auca, ANIC29034470 (1♀, ANIC) ; Rockwood Station , 23 km E. Condamine, S.E.Q ., -26.415218 150.917483, 18.viii.1984, D. Yeates, UQIC_ Reg 94767 (1♀, QM); Sheoak Ridge Nature Reserve near Julatten , -16.645258 145.403367, 9.xi.2014, C. Baker, M. Achatz, Malaise trap, CNC457380 (1♀, CNC) ; Six Mile Creek , W of Elimbah , -27.016667 152.933333, 15.ix.1985, G. and A. Daniels, GDCBReg_19570 (1♂, QM); Stanthorpe , -28.657928 151.930344, 14.x.1923, 613, ANIC29034846 (1♀, ANIC) ; 30.ix.1925 UQIC_ Reg 94760; CNC _ Diptera 253727 (2♂♂, QM; ANIC); Stockyard Creek , SW of Capalaba , -27.550000 153.166667, 25.ix.1982, G. Daniels, GDC- BReg_19669 (1♀, QM); Sunnybank , -27.578997 153.059006, 1950, I.M. Mackerras, Date is ambiguous reads ”3.9.50”, ANIC29034845 (1♂, ANIC) ; Tambourine Mountain , -27.923086 153.171717, 18.ix.1914, H. Hacker, QM _ REG._NO._T223168 (1♀, QM); Tibro Creek , -26.933333 152.950000, 26.viii.1960, F.A. Perkins, QM _ REG._ NO._T221266 (1♀, QM); Tibrogargan Ck., Glasshouse , -26.933333 152.950000, 29.viii.1958; 10.ix.1957, I.C. Yeo; F.A. Perkins, QM _ REG._NO._T221271-222172; UQIC_ Reg 94723 (1♂, 2♀♀, QM); Tin Can Beach , - 25.919297 153.003278, on Leptospermum sp., 21.viii.1957, F.A. Perkins, QM _ REG._NO._T221281 (1♂, QM); Toorbul Pt ., -27.033333 153.100000, 11.ix.1952, CNC _ Diptera 255054 (1♂, QM); Tully-Yabulu Hwy. 1 , -18.780303 146.126394, 29.viii.1986, E.A. Sugden, car net, Jeff _ Skevington _ Specimen 43813 (1♀, QM); Walten , -23.580000 149.130000, M. Taylor, ANIC2903460 (1♀, ANIC) ; Warrumbungle National Park ; Observatory , -31.274727 149.066153, Hilltop, 17.x.1997, J. Skevington, C. Lambkin, UQIC_ Reg 94753–94754 (2♂♂, QM); West Bluff Range Hictden Canyon via Biggenden , -25.600000 152.050000, 22.viii.1975, H. Frauca, ANIC29034999 (1♂, ANIC) ; West Claudie River, 4 km SW road junction, -12.733333 143.25, 1.xii.1986, G. Daniels & M.A. Schneider, malaise trap, UQIC_ Reg 94641 (1♀, QM); Wild Horse Mountain nr Beerburrum , -26.916667 153.000000, 12.iv.1986; 28.ix.1985, G. and A. Daniels; R. Eastwood, GDCBReg_19546; GDCBReg_19548–19549 (2♂♂, 1♀, GDCB ; Woombye, nr. Nambour , -26.666667 152.966667, 11–16.x.1965, D.H. Colless, ANIC29034528 (1♀, ANIC) ; Yarra- man, 2.2km SSE, -26.857000 151.994000, 28.iii–10.iv.2010, G. Monteith, malaise trap, QM _ REG._NO._T222150 (1♀, QM); nr. Lake Wabby, Fraser Island , -25.457644 153.129211, 9.viii.1981, M.A. Schneider, UQIC_ Reg 94758 (1♂, QM); rainforest nr. Rainbow beach , -25.897878 153.090378, rainforest, 14.xi.1982, G. and A. Daniels, GDC- BReg_19671 (1♀, QM) ; South Australia: Adelaide, Belair National Park , -35. 138.633333, hilltop, 4.x.1997, J.&A. Skevington, C. Lambkin, S. Winterton, Jeff _ Skevington _ Specimen 4413; UQIC_ Reg 94774 (2♂♂, QM); Ferguson, ANIC29034532 (1♀, ANIC) ; M[oun]t Lofty area near Adelaide, -35. 138.733333, 4.x.1997, J.&A. Skevington, C. Lambkin, S. Winterton, hand collected, Jeff _ Skevington _ Specimen 1246; Jeff _ Skevington _ Specimen 1247; Jeff _ Skevington _ Specimen 1248 (3♂♂, QM); Wyperfeld NP, East Lookout , -35.594192 142.112061, 21.x.2014, J.H., A.W., & A.M. Skevington, A.D. Young, A. Wyatt, & S. Namek, CNC384997 (1♂, CNC) ; Tasmania: 1 mile South of Scamander , -41.501533 148.248764, 18.i.1948, Key Carne & Kerr, ANIC29034935 (1♀, ANIC) ; 4 miles. E of Miena , -41.991797 146.780253, 26.i.1948, Key, Carne, & Kerr, ANIC29034500 (1♀, ANIC) ; 6 km S Deloraine , - 43.583333 146.650000, 12.i.1992, G. and A. Daniels, GDCBReg_19624 (1♀, GDCB); Advent Bay , 25–28.xii.1922, A. Tonnoir, ANIC29035779-80 ; ANIC29034543 ; CNC _ Diptera 253894; CNC_ Diptera 253895; CNC_ Diptera 253896; CNC_ Diptera 253897 (6♂♂, ANIC); Blackman Lagoon, Waterhouse , -40.916667 147.583333, on Banksia marginata , 18.i.1992, G. and A. Daniels, GDCBReg_19520 (1♂, GDCB); Cradle Mountain NP Lake Win- demere , -41.683478 145.953303, 1000m, M.P. van Zuijen, CNC _ Diptera 253785 (1♀, CNC); Cradle Mountain NP Pelion gap , -41.683478 145.953303, 1080m, 28.i.2006, M.P. van Zuijen; W. van Steenis, CNC _ Diptera 253786– 253787 (2♀♀, CNC); Cradle Mountain NP, Waterfall Valley , -41.683478 145.953303, 1000m, 26.i.2006, M.P. van Zuijen, CNC661784–661785 (2♀, WVSPC) ; Cradle Mountain National Park, Lake Windmere , -41.767125 145.956424, 1000m, 26.i.2006, M.P. van Zuijen, Jeff _ Skevington _ Specimen 43818 (1♀, QM); Cradle Mountain National Park, Pelion Gap , -41.864589 146.058961, 1080m, 28.i.2006, M.P. van Zuijen, Jeff _ Skevington _ Specimen 43819 (1♀, QM); Cradle Mountain National Park, Pencil Pine Creek , -41.683478 145.953303, 20.ii.1994, J. & A. Leask, AustralianMuseumK410097; AustralianMuseumK410479 (2♀♀, AM); Eaglehawk Neck , -43.019722 147.925000, G.H. Hardy, Date is ambiguous reads ”23 4.16/28”, AustralianMuseumK407605 (1♀, AM); Elizabeth Town , -41.466667 146.566667, 22.x.1997, A.I. Knight, QM _ REG._NO._T223230 (1♂, QM); Franklin-Gordon Wild River National Park, Donaghy’s Hill , -42.197778 145.931944, 11.i.2015; 7.i.1999, J.H., A.M. and A.W. Skevington; J. & A. Skevington, M. Matheison, ovipositing in huge fallen log, CNC457454 ; CNC457499 ; CNC451893 ; UQIC_ Reg 94556; UQIC_ Reg 94777; UQIC_ Reg 94780; UQIC_ Reg 94782 (5♂♂, 2♀♀, CNC, QM); Hobart , - 42.882139 147.327194, 29.xii.1955; i.1984, G.H. Hardy, AustralianMuseumK407686; ANIC29034843 (2♂♂, AM, ANIC) ; Lake St. Clair NP Cynthia Bay , -42.136267 146.230931, 740m, 22.i.2006, M.P. van Zuijen; W. van Steenis, CNC _ Diptera 253782 (1♀, CNC); Lake St. Clair NP Shadow Lake Circuit , -42.136267 146.230931, 800m; 850m, 22.i.2006, M.P. van Zuijen, CNC _ Diptera 253789–253790; CNC_ Diptera 253792 (3♀♀, WVSPC, MVZPC); Lake St. Clair Nat. Pk . 4 km N Derwent Bridge , -42.133333 146.183333, 29.i.1992, G. and A. Daniels, GDCBReg_ 19621, GDCBReg_19623 (1♂, 1♀, GDCB); Lake St. Clair National Park, Cynthia Bay , -42.125764 146.20603, 740m, 22.i.2006, M.P. van Zuijen; W. van Steenis, Jeff _ Skevington _ Specimen 43816–43817; Jeff _ Skevington _ Specimen 43823; Jeff _ Skevington _ Specimen 43825(4♀♀, MVZPC, WVSPC); Launceston , -41.433197 147.144094, 28.x.1916, F.M. Littler, ANIC29034860 (1♂, ANIC) ; Lispootah Dam, -42.318258 146.472692, 14.xi.1975, K.R. Norris, ANIC29034522 ; ANIC29034552 (1♂, 1♀, ANIC) ; Lyell Hwy: Franklin R nature trail, -42.215700 146.019800, on flowering Leptospermum , 11.i.2015, S.J. Grove, CNC _ Diptera 253819 (1♂, ANIC); Mole Creek , - 41.558628 146.404517, 4.ii.1916, G.H. Hardy, ANIC29034531 ; ANIC2903457 (2♀♀, ANIC) ; Mount William Na- tional Park; Summit of Mount William , -40.909722 148.186389, 23.xii.1998, J. &A. Skevington, UQIC_ Reg 94786; UQIC_ Reg 94789; UQIC_ Reg 94793 (3♂♂, QM); Mt Field NP Lake Belcher Track , -42.663558 146.649283, 1100m, 21.i.2006, M.P. van Zuijen; W. van Steenis, CNC _ Diptera 253793; Jeff _ Skevington _ Specimen 43812; Jeff _ Skevington _ Specimen 43814; Jeff _ Skevington _ Specimen 43824; CNC _ Diptera 253791; CNC_ Diptera 253780; CNC661778 ; CNC661783 (3♂♂, 6♀♀, WVSPC, MVZPC) ; Mt Field NP Tall Tree Nature Walk -42.663558 146.649283, 220m, 21.i.2006, W. van Steenis, CNC _ Diptera 254242 (1♀, WVSPC); Mt. Barrow Rd ., -41.345000 147.352778, 12–14.xii.1998, D. Yeates, S. Winterton, malaise trap, QM _ REG._NO._T221273–221275 (3♀♀, QM); Mt. Barrow , -41.366667 147.416667, 1000m, 23.i.1992, G. and A. Daniels, GDCBReg_19619 (1♀, QM); Mt. Wel- lington, Hobart , -42.916667 147.233333, 720m, 1.i.1992, G. Daniels, GDCBReg_19627–19628, GDCBReg_19632; GDCBReg_19635–19636;GDCBReg_19638–19639(2♂♂, 5♀♀, GDCB); Mt.Wellington , -42.890000 147.230000, G.H. Hardy, ANIC29034479 (1♀, ANIC) ; North of Corinna , -41.588514 145.189764, 12.i.2015, J.H., A.M. and A.W. Skevington, CNC457434 (1♀, CNC) ; Peilon Hut, 3km S Mt. Oakleigh , -41.500000 146.030000, 5–10.ii.1990, I.D. Naumann, CNC _ Diptera 253697 (1♀, ANIC); Strahan , -42.150000 145.316667, ii.1924, G.H. Hardy, ANIC29034519 ; CNC _ Diptera 253850–253860 (12♀♀, ANIC); Tarkine BushBlitz SG02 ridge NW of Lt Savage R, -41.472100 145.163200, 28.i.2015, S. Grove, CNC _ Diptera 253749 (1♀, TMAG); Tarkine BushBlitz SG 04 shru- bland adj quarry at Browns Plain , -41.610000 145.175500, 29.i.2015, S. Grove, CNC _ Diptera 253757; CNC_ Diptera 253755; CNC_ Diptera 253748 (2♂♂, 1♀, TMAG); Warra Long Term Ecological Research Site 106, Grid: 744282, LTIS8211, -43.1 146.683333, 17.xi–16.xii.1998, malaise trap, UQIC_ Reg 94603 (1♀, QM); Warra Long Term Research Site 59, Grid : 742279, LTIS8211, -43.1 146.683333, 14.i–12.ii.1998, malaise trap, UQIC_ Reg 94605 (1♀, QM) ; Victoria: 26 miles North Northeast, Orbost , -37.388111 148.591800, 396m, 6.xi.1969, I.F.B. Common, ANIC29034866 (1♂, ANIC) ; Ararat, -37.284600 142.931600, H.W. Davey, ANIC29034830 ; ANIC29035021 (1♂, 1♀, ANIC) ; Bettersby’s Camp, 15 km East of Nelson, lower Glenelg National Park , -38.030000 141.110000, 28.xi.1992, McAlpine, McEvey, Moulds,AustralianMuseumK407563 (1♀, AM); Croydon , -37.796000 145.281000, 17.xi.1909, E.M. Fulton, ANIC29034853 (1♂, ANIC) ; Grampians National Park , Reed’s Lookout , -37.148111 142.448261, 23.x.2014, J.H., A.M. and A.W. Skevington, CNC385028-385030 (2♂♂, 1♀, CNC) ; Melbourne, - 37.813319 144.963336, G.H. Hardy, ANIC29034492 (1♀, ANIC) ; Mt. Buffalo NP Dickson Falls , -36.731325 146.784042, 1440m, 29.i.2006, W. van Steenis, CNC _ Diptera 253781; CNC_ Diptera 253763, CNC_ Diptera 253776; CNC661775 ; Jeff _ Skevington _ Specimen 43815; Jeff _ Skevington _ Specimen 43820–43821 (2♂♂, 5♀♀, WVSPC); Mt. Dom Dom , -37.590278 145.657778, 22.x.1961, D.H. Colless, ANIC29034468 (1♀, ANIC) ; Princess Margaret, Rose Caves, 8 km North Northwest of Nelson, -37.59 140.58, 29.xi.1992, McAlpine, McEvey, Moulds, sweeping, AustralianMuseumK407565 (1♂, AM); Urquhart Bluff , -38.436336 144.130617, 19.xi.1975, K.R. Norris, ANIC29034879 (1♂, ANIC) ; Warburton District , -37.752539 145.694839, 9.xii.1900, ANIC29034539 (1♀, ANIC) ; Wyperfield National Park , -35.588747 142.051722, 5.xi.1966, I.F.B. Common & M.S. Upton, ANIC29035010 (1♂, ANIC) ; Young’s Ck 12 km N of Orbost Bohang Hwy. , -37.609489 148.466761, 8.xi.1976, Colless & Liepa, USNM _ ENT249834 ; CNC _ Diptera 253872 (2♂♂, ANIC) ; Western Australia: 2 km North Northwest of Crossman , - 32.750000 116.566667, 24.ix.1981, I.D. Naumann, J.C. Cardale, ANIC29034834 (1♂, ANIC) ; 24 mi. NW of Wal- pole, -34.681372 116.514911, 7.x.1970, D.H. Colless, light trap, ANIC29034546 ; ANIC29035008 (2♂♂, ANIC) ; 8 km S Permberton , -34.483333 116.000000, 17.i.1987, G. and A. Daniels, GDCBReg_19730 (1♀, GDCB) ; 9 km South Southwest of Bindoon , -31.448614 116.064672, 25.ix.1981, J.C. Cardale, light trap, ANIC29034996 (1♂, ANIC) ; Bluff Knoll Road, Stirling National Park , -34.399117 117.962769, 12.x.1970, D.H. Colless, ANIC29034924 (1♀, ANIC) ; Circular Pool, -34.943903 116.799967, 9.x.1970, D.H. Colless, ANIC29034510 (1♀, ANIC) ; Cran- more Park, -30.602581 116.297339, 25.ix.1932, ANIC29034472 ; ANIC29034483 (2♀♀, ANIC) ; Crowea S/F Pem- berton, -34.533467 116.156211, 15.xi.1978, S.J. Curry, ANIC29034524 (1♂, ANIC) ; Crystal Springs, 7 miles West of Walpole , -34.903975 116.621292, on leaves of Xanthorrhoea , 14.xii.1970, G.A. Holloway & H. Hughes, AustralianMuseumK408008 (1♀, AM); D’Entrecastaux Pk , -34.775028 116.163278, ex. coastal heath, flowering, 17.xi.2008, S.D. Gaimari & S.L. Winterton, CNC661771 ; CNC661774 (1♂, 1♀, CSCA) ; Dwellingup, -33.000000 116.050000, 3.x.1981, I.D. Naumann, J.C. Cardale, ANIC29034494 (1♀, ANIC) ; Hovea, -31.887233 116.115558, 29.ix.1935, K.R. Norris, ANIC29034447 (1♀, ANIC) ; Leeuwin-Naturalists Natl Park, -34.050778 115.018167, ex. coastal heath, flowering, 16.xi.2008, S.D. Gaimari & S.L. Winterton, CNC661773 (1♂, CSCA) ; Leewin Naturaliste NP: Boranup Beach Road , -34.159444 115.028889, 50m, coastal heath, 6.xi.2003, C. Lambkin, J. Recsei, A. Zwick, ANIC29035756 (1♀, ANIC) ; Mount Magog , Stirling Ranges , -34.399117 117.962769, 7–8.xii.1970, G.A. Hollo- way, malaise trap, AustralianMuseumK410145 (1♀, AM); Mundaring , -31.897006 116.171136, 23.viii.1926, E.W. Ferguson, ANIC29034448 ; ANIC29034992 (1♂, 1♀, ANIC) ; Nannup, -33.982000 115.765000, 12.xi.1958, E.F. Riek, ANIC29034544 (1♂, ANIC) ; Pemberton, -34.440000 116.050000, 29.xi.1936, K.R. Norris, ANIC29034874 (1♂, ANIC) ; S.W. District, ix.1938, E.H. Kipps, ANIC29034477 (1♀, ANIC) ; Stirling Nature Park, 49 mile post, - 34.399117 117.962769, 12.x.1970, D.H. Colless, ANIC29034503 ; CNC _ Diptera 253849 (2♀♀, ANIC); Stirling Nature Park , 54 mile post , -34.399117 117.962769, 12.x.1970, D.H. Colless, ANIC29034502 (1♀, ANIC) ; Stirling Range Drive, Stirling Range National Park , -34.399117 117.962769, 23.ix.1981, I.D. Naumann, J.C. Cardale, ANIC29034995 (1♂, ANIC) ; Stirling Range National Park , Bluff Knoll, Hilltop , -34.377500 118.252222, 28.xii.1999, J.&A. Skevington, QM _ REG._NO._T221265 (1♂, QM); Stirling Range, Mount Abbey , -34.349992 117.950172, 20.x.1977, Signe Holm leg., JeffSkevingtonSpecimen#31419 (1♂, ZMUC); Stirling Range, Mount Hassel , -34.377776 118.068697, 25.x.1977, Signe Holm leg., JeffSkevingtonSpecimen#31420 (1♂, ZMUC); Stir- ling Range, Toolbrunup , -34.382569 118.049252, 20.x.1977, Signe Holm leg., JeffSkevingtonSpecimen#31418 (1♂, ZMUC); Toolbrunup Camp, Stirling Range NP , -34.399117 117.962769, 22.ix.1981, J.C. Cardale, ANIC29034525 (1♂, ANIC) ; Treen Brook, 5 miles West of Pemberton, -34.449886 115.950942, 19.xii.1970, G. Williams & H. Hughes, Malaise trap, AustralianMuseumK410505 (1♀, AM); Warren River , 6 miles Southeast of Pemberton, -34.511628 116.114703, 17.i.1971, G.A. Holloway & H. Hughes, AustralianMuseumK410130 (1♂, AM); near Warren River , 10 km South of Pemberton , -34.534142 116.046947, 27–29.ix.1987, D.K. McAlpine, AustralianMuseumK407956 (1♀, AM) .

Discussion: Psilota tristis doubtlessly encompasses multiple species as described here. Initial DNA barcode analysis indicate that at least 23 species are represented. Further DNA barcoding and male genitalia dissection is required to adequately separate the full range of specimens into species and will be discussed in a future revision. Furthermore, there are several other species of Psilota described from Australia that may or may not be synonyms of P. tristis , including Psilota hirta , Psilota shannoni , and Psilota queenslandica (see ‘species not here recognized’ below). Finally, the type specimens of P. tristis , P. queenslandica , and P. hirta were deposited in Venezuela and were unavailable for study at the time of this publication. Because it is impossible to tell what name should properly be assigned to different Barcode identities without a full revision of this complex, these species have not been treated individually.

Figures

FIGURE 35. P. tristis Klocker. A) male dorsal habitus, B) male anterolateral face, C) male lateral habitus. Scale bar =1 mm.

FIGURE 64. Known range of A) P. spinifemur sp. nov.; B) P. tristis Klocker.