Schizotergitius altajicus Loksa, 1978

Dyachkov, Yu. V., Farzalieva, G. Sh. & Dányi, L., 2021, On the centipede genus Schizotergitius Verhoeff, 1930, with a redescription of Schizotergitius altajicus Loksa, 1978 and a key to the genera of the family Lithobiidae in Central Asia (Chilopoda: Lithobiomorpha), Russian Entomological Journal 30 (3), pp. 361-371 : 364-371

Schizotergitius altajicus Loksa, 1978

Figs 1–53 .

Schizotergitius altajicus Loksa, 1978: 116–118 , figs 10–19.

Schizotergitius altajicus forma gracile (sic!) Loksa, 1978: 118.

Schizotergitius altajicus — Eason, 1986a: 190.

Schizotergitius altajicus — Dyachkov, 2017: 35 .

Schizotergitius — Farzalieva et al., 2017: 124, table 1.

DIAGNOSIS. The species differs from S. longiventris by the smaller size (12–21 mm vs. 22.5– 25 mm in S. longiventris ), body colouration (yellow or brownish yellow vs. a greybrown body with a darker head in S. longiventris ), the presence of DCa on 15 legs (vs. absent in S. longiventris ), as well as the number of dorsal spines on 13P (three dorsal spines vs. a single dorsal spine on 13P in S. longiventris ).

BRIEF DESCRIPTION. A medium-sized (12–21 mm) Lithobiinae with 5–6 ocelli in a single broken row, Tömösváry´s organ smaller than the nearest ocelli; usually 20 antennal articles (rarely 17–22); forcipular coxosternite with 2+2 acute teeth and setiform porodonts; TT 3, 5, 8, 10, 12 with posterior median notches, SS elongated (excluding genital S); legs 1–13 with functionally biarticulate tarsi; legs 15 without accessory spines; DCa present on legs (11)12–15; number of coxal pores, 2–4 on legs 12–15; ♂ 15F (sometimes 15P) showing more or less evident, indistinct, wide, dorsal and/or dorsolateral sulci, 15Ti with a dorsal or dorsolateral sulcus and a dark blot in distodorsal part covered with numerous setae; ♂ gonopod 1-segmented with 2–4 setae; ♀ gonopod with 2+2 gonopodal spurs and a simple, curved and pointed claw, 1 st segment without dorsal setae (rarely with 1), 2 nd segment with a small, translucent, distodorsal outgrowth shifted mesad ( Figs 28 –30) and 4–6 dorsal setae, 3 rd segment with 2–4 dorsal setae (Figs 31–33, 36–37).

MATERIAL. Type material: paratypes: 12 ♂♂, 10 ♀♀ [vs. 11 ♂♂, 11 ♀♀, 4 juv. in the original description] ( HNHM chilo-7558) (one ♂ with postpedal segments, forcipules, the left ocellar field, T7 , left 1 st and right 14–15 th legs missing, probably removed and mounted on slides which could have served for Loksa´s [1978] illustrations, but no such slides could be relocated], [ Mongolia, Govi-Altai Aimag, Mt. Range, 10 km S Somon Cogt, ca. N45°20´, E96°38´] “ Gobi Altaj Gebirge, 10 km S. von Somon Cogt, am Pass, 2500 m, 27.VI.1966 (Nr. 589)”; Schizotergitius altajicus f. gracile Loksa, 1978, types: 3 ♀♀ [♀ holotype, 1 ♂ and 1 ♀ paratype, according to the original description, but no holotype selected and labeled in the material] ( HNHM chilo-7559), [ Mongolia, Govi-Altai Aimag, between Lake Beger-Nuur and Somon Beger, ca. N45°44´, E97°09´] “zwischen dem See Beger Nuur und Somon Beger , 1400 m, 25–26.VI.1966 (Nr.579)”.

Other material: Mongolia: 4 ♀♀ ( ASU No. 295), Bayan-Ölgii Aimag, Buratyn-Davaa Pass , N48°28´, E90°27´, mountain stony steppe, 2600 m, 24. VI.2015, coll. AF, OB, DN ; 8 ♂♂ ( ASU No. 320), 24. VI 2015, coll. AF, OB, DN ; 1 ♂, 1 ♀ ( ASU No. 41), Khovd Aimag, Sutai-Uul Mt. Range, Mt Zamlyn-Bogt , scree, N46°39´, E93°42´, 3810–3860 m, coll. AF ; 2 ♂♂, 1 ♀ ( ASU No. 42), Bumbat- Khayrhan-Bogt Mt. Range, Mt Degnuul , stony alpine meadow, N47°12´, E93°08´, 3000–3200 m a.s.l., 24. VI.2017, coll. AF ; 1 ♂, 2 ♀♀ ( ASU No. 296), Baitag-Bogd-Uul Mt. Range, Gushoot-Shyneetiyn-Gol River Valley , N45°16´, E91°04´, 1900 m, 23. V.2015, coll. VD, AK ; 2 ♂♂, 1 ♀ ( PSU No. 685), N45°15´, E91°03´, 1900 m a.s.l., 24. V.2015, coll. RY ; 2 ♂♂ ( ASU No. 293), Dzhargalant-Khairkhan Mt. Range, near source of Ar-Shaatyn-Gol River , mountain stony tundra, N47°43´, E92°29´, 3300–3500 m a.s.l., 25. VI.2015, coll. AF ; 1 ♂ ( ASU No. 294), Ikh-Khavtag-Uul Mt. Range, near Zhugentiyn Spring , N45°04´, E92°13´, mountain stony steppe with rocks, 2050– 2300 m a.s.l., 30. VI –2.VII.2015, coll. AF ; 1 ♂ ( PSU No. 668), Ikh-Khavtag-Uul Mt. Range, near Yargaytiyn-Sayr Ravine , mountain stony steppe with rocks, N45°04´, E92°13´, 1900–2150 m a.s.l., 30– 31. V.2015, coll. AF ; 4 ♂♂, 1 ♀ ( HNHM chilo-7686), Govi-Altai Aimag, Cakhir-Khaalgyn-Nuruu Mt. Range , 2–5 km SSE Dotiyn-Davaa Pass, mountain stony tundra with rocks and scree, N45°25´, E97°05´— N45°24´, E97°07´, 3300–3660 m a.s.l., 04.VII.2017, coll. AF ; 1 ♂ ( HNHM chilo-7688), 04.VII.2017, coll. AF ; 1 ♀ ( HNHM chilo-7687), foothill of Cakhir-Khaalgyn-Nuruu Mt. Range, near Mt Dotiyn-Davaa , pass, stony alpine meadow, N45°25´, E97°05´, 2900– 3000 m a.s.l., 04–05.VII.2017, coll. AF ; 4 ♂♂, 2 ♀♀ ( ZMMU Rc 7892), Khasagtiyn-Nuruu Mt. Range, vicinity of Mt Khoit-Bogd- Uul , mountain stony tundra with rocks and scree, N46°47´, E95°50´, 3400–3500 m a.s.l., 08.VII.2017, coll. AF .

REDESCRIPTION. ♂ paratypes. Body 12–17 mm long, colouration in alcohol yellow ( Fig. 2 ), cephalic plate slightly brighter; head and body slightly punctate. Head a little broad- er than T1 ( Fig. 41 ) (ca. 1:0.95).

Antennae with 20 articles (1 ♂ with 18+20 ones, 2 ♂♂ with 19+20), covered with dense, short, dark and erect sensilla. Length to breadth ratio of terminal article, 1:0.4 ( Fig. 44 ). Antennae extending back to ca. T4.

Ocelli: 5–6 on each side arranged in a single broken row inside a contoured area ( Figs 13, 14 ). Tömösváry´s organ smaller than nearest ocelli, rounded.

Coxosternum with 2+2 acute teeth and setiform porodonts, median diastema V-shaped. Shoulders very gently sloping behind porodonts.

Tergites: almost smooth, with short and sparse setae, without posterior triangular projections. TT 3, 5, 8, 10, and 12 elongated (Table 3); posterior median notch developed on TT 3, 5, 8, 10 and 12 (caudal part of each of these TT with a well-developed, light, short suture dividing it into two lobes) ( Figs 4–8 , 51 ); T15 indistinct. T1 breadth ca. 1.2 mm, T8 breadth ca. 1.45 mm. All microtergites (TT 2, 4, 6, 9, 11, 13) and T7 with one median and two paramedian, short, translucent sutures at posterior margin ( Fig. 50 ). Caudal margin of Tim slightly sinuate, length to breadth ratio, 1:0.94 ( Fig. 42 ). Caudal part of 14–15 th sternites and coxae densely covered with short setae ( Fig. 45 ).

Sternites elongate (except for genital S); posterior margin of SS 1–14 with short sutures (best expressed on SS 2–12).

All Tsi functionally biarticulate, with Ts1 about 2 times longer than Ts2. Tsi 1–13 each with a ventral row of dark setae especially well-developed on midbody legs. Legs 1–14 each with a posterior accessory spine, legs 1–13 each with a large, pretarsal, anteroventral spine (Fig. 35). Legs 15 without accessory spines ( Fig. 21 ), but with a very small anteroventral spine (interpreted as an accessory spine in the original description). P, F, Ti of legs 15 slightly incrassate and flattened dorsally ( Figs 19–20 ). Legs 14 without visible secondary sexual characters.

15F covered with sparse minute setae and several small, oval, dark pores ( Figs 26, 27 ). Distodorsal part of 15Ti flattened and with a long, median, dark blot formed by numerous small pores, as well as with numerous dense setae located on and around the dark blot ( Figs 22, 23, 25 ); 15F and 15Ti with more or less evident, shallow, wide, indistinct dorsal and/or dorsolateral sulci ( Figs 20, 26 ). Leg plectrotaxy as in Table 1.

DCa developed starting with legs (11)12. Coxal pores small, rounded, separated from one another by a distance about 2–4 times greater than their own diameter, formula 3,2(3),2(3),2(3) ( Fig. 45 ). Gonopod ( Fig. 24 ) 1-segmented, with 3–4 very thin and light setae placed at gonopodal middle.

♂ variations (non-type material).

Most characteristics as in paratypes, except as follows. Body length 13–21 mm, colouration in alcohol always brownish yellow ( Fig. 3 ). Antennae with 20–22 segments

Table 1. Plectrotaxy of Schizotergitius altajicus Loksa, 1978 (paratypes). Brackets indicate variable spines. Таблица 1. Распределение Шипов на ногах у Schizotergitius altajicus Loksa, 1978 (паратипы). Скобки обоЗначают вариабельные Шипы. <

Figs 30–40. Schizotergitius altajicus Loksa, 1978 (♀♀ paratypes: 30–33, 35; ♀ non-type: 34, 36–40 — ASU No. 295): 30, 31 — gonopods, ventral and ventrolateral views, respectively; 32 — right gonopod, dorsolateral view; 33, 34 — left gonopod, dorsal and lateral views, respectively; 35 — pretarsus of leg 14; 36 — fragment of right gonopod, ventrolateral view; 37 — left gonopod, dorsolateral view; 38 — right gonopod segments 2 and 3, lateral view; 39, 40 — right gonopod, distal and distodorsal views, respectively. Abbreviations: as — posterior accessory spine; do — distodorsal outgrowth, s — anteroventral spine. Scale: 0.2 mm (30, 31), 0.1 mm (32–40).

Рис. 30–40. Schizotergitius altajicus Loksa, 1978 (паратипы ♀♀: 30–33, 35; нетиповаЯ ♀: 34, 36–40 — ASU No. 295): 30, 31 — гоноподы, соответственно сниЗу и одновременно сниЗу и сбоку; 32 — правый гонопод, одновременно сверху и сбоку; 33, 34 — левый гонопод, соответственно сверху и сбоку; 35 — коготь ноги 14; 36 — фрагмент правого гонопода, одновременно сниЗу и сбоку; 37 — левый гонопод, одновременно сверху и сбоку; 38 — сегменты 2 и 3 правого гонопода, сбоку; 39, 40 — правый гонопод дистально и дистодорсально. ОбоЗначениЯ: as — дополнительный коготок; do — дистальный верхний вырост, s — передне- нижнЯЯ Щетинка. МасШтаб: 0,2 мм (30, 31), 0,1 мм (32–40).

(19+ 20 in 1 specimen), terminal segment of antennae as in Fig. 43 .

Dental margin of forcipular coxosternite as in Fig. 47 .

Sides of labrum with well-expressed fringes of numerous thin bristles; a pair of setae projecting across labral midpiece ( Fig. 15 ). Gnathal edge of mandible with numerous pulvilli, 4 pairs of well-developed teeth and 10–12 rather thick and serrate aciculae ( Figs 9–10 ). First maxillae: edge with more than 20 plumose bristles and simple setae as well ( Figs 16– 18 ). Second maxillary telopodite ( Figs 11, 12 ) with bristles, both simple and plumose, at tip (more than 25 plumose bristles on internal side).

Hind margin of Tim strongly sinuate, and posterior margins of lobes slightly deflexed down.

15P rarely with indistinct dorsal and/or dorsolateral sulci; 15F dorsally with 3–7 oval pores ( Figs 48, 49 ). Dark blot on 15Ti consisting of numerous, densely located, small pores ( Figs 28, 29 ). Leg plectrotaxy as in Table 2.

SS 14 and 15, coxae 13–15 ( Fig. 45 ), as well as ventral sides of P, F and Ti 12 and 13 covered with very small and dense setae in rear parts and, in some cases, even all along the articles ( Fig. 54 ). Genital sternite short and broad, covered with long setae.

Each gonopod with 3 (rarely 2) setae.

♀ paratypes. Most characteristics as in ♂ paratypes, but body (11–16 mm) and TT shorter than in ♂ (Table 3). T1 breadth ca. 1.01 mm, T8 breadth ca. 1.29 mm. Coxosternum usually with 2+2 acute teeth (1 specimen with 2+1, another one with 2+3 ones). Formula of coxal pores, 3,3,3,3.

Gonopod with 2+2 acute spurs (Figs 30, 31) and a simple, pointed and curved claw; 1 st segment without dorsal setae, but with 1–3 short and 3 longer medial setae, 2 nd segment with a small, translucent, distodorsal outgrowth shifted mesad and 4–6 dorsal setae (vs. 7 in the holotype in the original description, but, according to the illustrations [ Loksa, 1978: figs 17, 19], a lateral seta similar in appearance was also counted), 3 rd segment with 2–3 dorsal setae (Figs 32, 33).

♀ variations (non-type material).

Body 12–20 mm long, antennae usually with 20 antennomeres (1 ♀ with 17+19).

Table 2. Plectrotaxy of Schizotergitius altajicus Loksa, 1978 (non-types). Brackets indicate variable spines. Таблица 2. Распределение Шипов на ногах у Schizotergitius altajicus Loksa, 1978 (нетиповой материал). Скобки обоЗначают вариабельные Шипы. Table 3. Average lengths and breadths of tergites of ♂♂ and ♀♀ in Schizotergitius altajicus Loksa, 1978 (paratypes).

Таблица 3. Средние ЗначениЯ длины и Ширины тергитов у самок и самцов Schizotergitius altajicus Loksa, 1978 (паратипы).

Forcipular coxosternum as in Fig. 46 .

Posterior margin of SS 1–14 with short sutures ( Figs 52, 53 ). Formula of coxal pores, 3(4),3,3,2(4).

Gonopods: each with 2+2 acute spurs (Fig. 34) and a simple, pointed and curved claw (Figs 36–40) with irregular edges (often broken off); 1 st segment mostly without dorsal setae (1 dorsal seta on one or both gonopods presents (Fig. 37) in 3 of 4 ♀♀ from Bayan-Ölgii Aimag (ASU No. 295)), with up to 6 medial setae, 2 nd with 4–6, 3 rd with 3–4 dorsal setae in a single row; 2 nd segment with a small, distodorsal and transparent outgrowth shifted mesad.

DISTRIBUTION. Western Mongolia: Bayan-Ölgii, Khovd and Govi-Altai Aimags [ Loksa, 1978; Dyachkov, 2017], summarized in Fig. 1 .

REMARKS. Three specimens of S. altajicus f. gracile (recte: gracilis ) are labeled as 1 ♂ and 2 ♀♀ in the original description [♀ holotype, 1 ♂ and 1 ♀ paratype], but actually all three are ♀♀: one ♀ has postpedal segments damaged, and legs 15 are devoid of the secondary sexual characters of the ♂. No holotype is labeled as such in the vial.

An important feature of S. altajicus is its remarkable sexual dimorphism. ♂♂ show the body and macrotergites more strongly elongated than ♀♀ do, but this character is not unique, as the same pattern is observed in the Asiatic Lithobius giganteus Sseliwanoff, 1881 [ Eason, 1986a]. This similarity between S. altajicus and L. giganteus was noted by Eason [1986a] who mentioned tergal modifications in some specimens of L. giganteus ( Figs 55–57 ), as well as “tergite 12 in the fragmented female of [ Lithobius ] mongolicus [now L. giganteus ] is exactly the same as that of tergite 5 in S. altajicus ”. However, S. altajicus differs from L. giganteus in the shorter body, as well as the presence of the well-developed secondary sexual modifications of ♂ legs 15: 15F has a row of 3–7 oval and dark pores, 15Ti shows a dark, distodorsal, elongate blot (consisting of numerous pores) covered with dense and short setae. Similarly dense setae are also present in another Asiatic genus, Disphaerobius Attems, 1926 , but these are located on a rounded outgrowth of Ti, while a blot is absent [ Farzalieva et al., 2017: fig. 5].

The following eight genera of Lithobiidae are presently known to occur in Central Asia: Bothropolys Wood, 1862 (Ethopolyinae Chamberlin, 1915), Australobius Chamberlin, 1920 , Disphaerobius Attems, 1926 , Hessebius Verhoeff, 1941 , Lithobius Leach, 1814 , Schizotergitius Verhoeff, 1930 , and Validifemur Ma, Song & Zhu, 2007 ( Lithobiinae Verhoeff, 1907) [ Loksa, 1978; Zalesskaja, 1978; Eason, 1997; Ma et al., 2007, 2014; Farzalieva et al., 2017; Ganske et al., 2020]. Now that the identity of Schizotergitius has become clarified, we can propose the following key for their separation.

A KEY TO THE GENERA OF LITHOBIIDAE REPORTED FROM CENTRAL ASIA

1 Coxal pores scattered ................................. Bothropolys

— Coxal pores arranged in a single row ( Fig. 45 ) ........... 2

2 Tömösváry´s organ located below interior row of ocelli ................................................................... Australobius

— Tömösváry´s organ located level with interior row of ocelli ( Figs 13, 14 ) ....................................................... 3

3 Tergites 3, 5, 8, 10, 12 with posterior median notches ( Figs 4–8 , 51 ) ......................................... Schizotergitius

— Tergites without median notches ................................. 4

4 ♂ .................................................................................. 5

— ♀ .................................................................................... 7

5 Tergites 12–14 (or some of them) modified: serrate and sometimes broadened ............................ Disphaerobius

— Tergites neither serrate nor broadened ........................ 6

6 Leg-pair 15 without falciform spines ............................. ................................................ Hessebius and Lithobius

— Leg-pair 15 with strong falciform spines ... Validifemur

7 Second ♀ gonopodal article with neither a distodorsal protuberance nor an outgrowth ................................... 8

— Second ♀ gonopodal article with a distodorsal protuberance or outgrowth ........................................................ 9

8 Tarsus of legs 1–13 without any marking on dorsal side, but with a trace of bipartite division on ventral side ..... ..................................................................... Validifemur

— Tarsus of legs 1–13 with or without a bipartite division ......................................................................... Lithobius

9 Second ♀ gonopodal article with a more or less developed protuberance, often with setae or spines ....... Hessebius

— Second ♀ gonopodal article without such protuberance, but with a small, oval, distodorsal outgrowth directed mesad, always with neither setae nor spines ........ Disphaerobius

Acknowledgements. We are most grateful to S.I. Golovatch (Moscow) who kindly checked the English of an advanced draft. We are highly obliged to A.A. Fomichev who donated us his material for study. The first author is grateful to R.V. Yakovlev who organized his trip to Budapest, and to O.V. Fast, L.A. Khvorova, and A.V. Vaganov (all Barnaul, Russia) who kindly provided FAST software for taking the measurements.

Competing interests. The authors declare no competing interests.

Figures

Fig. 1. Distribution of Schizotergitius altajicus Loksa, 1978: the type locality (asterisk), the type locality of S. altajicus f. gracilis Loksa, 1978 (triangle), non-type material (circles). Рис. 1. Распространение Schizotergitius altajicus Loksa, 1978: типовое местонахождение (ЗвеЗдочка), типовое местонахождение S. altajicus f. gracilis Loksa, 1978 (треугольник), нетиповой материал (круг).

Figs 2–8. Schizotergitius altajicus Loksa, 1978 (2, 4–8 — ♂ paratypes, 3 — ♂ non-type, HNHM chilo-7688), dorsal view: 2, 3 — habitus; 4 — tergite 3; 5 — tergite 5; 6 — tergite 8; 7 — tergite 10; 8 — tergite 12. Abbreviation: mn — median notch. Scale: 2 mm (2, 3), 0.5 mm (4–8). Рис. 2–8. Schizotergitius altajicus Loksa, 1978 (2, 4–8 — паратипы ♂♂, 3 — нетиповой ♂, HNHM chilo-7688), сверху: 2, 3 — обЩий вид; 4 — тергит 3; 5 — тергит 5; 6 — тергит 8; 7 — тергит 10; 8 — тергит 12. ОбоЗначение: mn — срединнаЯ бороЗдка. МасШтаб: 2 мм (2, 3), 0,5 мм (4–8).

Figs 9–18. Schizotergitius altajicus Loksa, 1978 (♂ paratypes: 13, 14; non-types: 9, 11, 12, 15–18 — PSU No. 668, 10 — ASU No. 295): 9 — mandible, anterodorsal view; 10 — aciculae of mandible, anterodorsal view; 11 — right part of second maxillae and its apical part, ventral view; 12 — apical part of second maxillae, dorsal view; 13–14 — ocelli and Tömösváry´s organ, lateral view; 15 — central and left parts of labrum, ventral view; 16 — apical part of first maxillae telopodite, ventral view; 17 — coxal projection of first maxillae, ventral view; 18 — terminal article and coxal projection of the first maxillae, ventral view. Scale: 0.1 mm (11–12, 15–18), 0.5 mm (9), 0.05 mm (10). Рис. 9–18. Schizotergitius altajicus Loksa, 1978 (паратипы, самцы: 13, 14; нетиповой материал: 9, 11, 12, 15–18 — PSU No. 668, 10 — ASU No. 295): 9 — мандибула, одновременно спереди и сверху; 10 — мандибулЯрные ацикулы, одновременно спереди и сверху; 11 — праваЯ часть вторых максилл и их апикальнаЯ часть, сниЗу; 12 — верШиннаЯ часть вторых максилл, сверху; 13–14 — глаЗки и орган ТемеШвари, сбоку; 15 — центральнаЯ и леваЯ части верхней губы, сниЗу; 16 — верШиннаЯ часть телоподита первых максилл, сниЗу; 17 — коксальный отросток первых максилл, сниЗу; 18 — верШиннаЯ часть телоподита и коксальный отросток первых максилл, сниЗу. МасШтаб: 0,1 мм (11–12, 15–18), 0.5 мм (9), 0,05 мм (10).

Figs 19–29. Schizotergitius altajicus Loksa, 1978 (male; paratypes: 19–27; nontype: 28, 29 — PSU No. 668): 19 — leg 15, dorsal view; 20 — femur and tibia of legs 15, dorsal view; 21 — claw of leg 15, lateral view; 22 — tibia 15, dorsal view; 23 — same, meso-dorsal view; 24 — gonopod, ventral view; 25 — distal part of tibia 15, mesodorsal view; 26 — femur 15, dorsal view; 27 — same, mesodorsal view; 28 — distal part of tibia 15, dorsal view; 29 — blot on tibia 15, dorsal view. Abbreviations: ds — dorsal sulcus, dls — dorsolateral sulcus, p — pore. Scale: 1 mm (19), 0.5 mm (20, 22, 23, 25–28), 0.2 mm (29), 0.1 mm (21, 24). Рис. 19–29. Schizotergitius altajicus Loksa, 1978 (♂♂; паратипы: 19–27; нетиповой материал: 28, 29 — PSU No. 668): 19 — нога 15, дорсально; 20 — бедро и голень ноги 15, дорсально; 21 — коготок ноги 15, латерально; 22, 23 — голень ноги 15, соответственно сверху и одновременно иЗнутри и сверху; 24 — гонопод, вентрально; 25 — дистальнаЯ часть голени 15, меЗо-дорсально; 26 — бедро 15, дорсально; 27 — то же, меЗо-дорсально; 28 — дистальнаЯ часть голени 15, дорсально; 29 — пЯтно на голени 15, дорсально. ОбоЗначениЯ: ds — дорсальнаЯ бороЗдка, dls — верхнебоковаЯ бороЗдка, p — пора. МасШтаб: 1 мм (19), 0.5 мм (20, 22, 23, 25–28), 0.2 мм (29), 0.1 мм (21, 24).

Figs 41–49. Schizotergitius altajicus Loksa, 1978 (paratypes: 41, 42, 44, 45; non-types: 43, 46, 48 — ASU No. 295, 47, 49 — PSU No. 668): 41 — front body fragment of ♂, dorsal view; 42 — intermediate tergite of ♂, dorsal view; 43, 44 — fragment of ♂ antenna, lateral and dorsal views, respectively; 45 — rear body fragment of ♂, ventral view; 46 — ♀ forcipule, ventral view; 47 — dental margin of ♂ forcipular coxosternite, ventral view; 48, 49 — ♂ femur 15, dorsal view. Abbreviation: p — pore. Scale: 1 mm (49), 0.5 mm (41, 42, 45, 46, 48), 0.1 m (43, 44), 0.3 mm (47). Рис. 41–49. Schizotergitius altajicus Loksa, 1978 (паратипы: 41, 42, 44, 45; нетиповой материал: 43, 46, 48 — ASU No. 295, 47, 49 — PSU No. 668): 41 — переднЯЯ часть тела ♂, сверху; 42 — промежуточный тергит ♂, сверху; 43, 44 — фрагмент антенны ♂, соответственно сбоку и сверху; 45 — ЗаднЯЯ часть тела ♂, сниЗу; 46 — ногочелюсть ♀, сниЗу; 47 — Зубной край коксостернита ногочелюсти ♂, сниЗу; 48, 49 — бедро 15 у ♂, сверху. ОбоЗначение: p — пора. МасШтаб: 1 мм (49), 0,5 мм (41, 42, 45, 46, 48), 0,1 мм (43, 44), 0,3 мм (47).

Figs 50–57. Schizotergitius altajicus Loksa, 1978 (50–54, paratypes) and Lithobius giganteus Sseliwanoff, 1881 (55–57, ♂, ASU No. 318; Mongolia, Khovd Aimag; dorsal view): 50 — ♂ tergite 2, dorsal view; 51 — rear part of ♂ tergite 5, dorsal view; 52, 53 — ♀ sternites 7 and 8, ventral view; 54 — prefemur and femur of ♂ leg 13, ventrolateral view; 55 — rear part of tergite 12; 56 — tergites 4 and 5; 57 — tergites 9 and 10. Abbreviations: n — notch, mn — median notch. Scale: 1 mm (56, 57), 0.5 mm (50–53, 55), 0.2 mm (54). Рис. 50–57. Schizotergitius altajicus Loksa, 1978 (50–54, паратипы) и Lithobius giganteus Sseliwanoff, 1881 (55–57, ♂, ASU No. 318; МонголиЯ, Ховд Аймак; сверху): 50 — тергит 2 у ♂, сверху; 51 — ЗаднЯЯ часть тергита 5 у ♂, сверху; 52, 53 — стерниты 7 и 8 у ♀, сниЗу; 54 — предбедро и бедро 13-й пары у ♂, одновременно сниЗу и сбоку; 55 — ЗаднЯЯ часть тергита 12; 56 — тергиты 4 и 5; 57 — тергиты 9 и 10. ОбоЗначениЯ: n — бороЗдка, mn — срединнаЯ бороЗдка. МасШтаб: 1 мм (56, 57), 0,5 мм (50–53, 55), 0,2 мм (54).