Tremapterus (Nyanja) Distant

Andrew Hamilton, K. G., 2015, A new tribe and species of Clastopterinae (Hemiptera: Cercopoidea: Clastopteridae) from Africa, Asia and North America, Zootaxa 3946 (2), pp. 151-189 : 176

Tremapterus (Nyanja) Distant , stat.nov.

Nyanja Distant, 1908 b:224 . Type-species by monotypy: N. muiri Distant, 1908 b .

Diagnosis. Face ivory or yellow to redbrown with frons transversely banded with black between antennal ledges (as in Abbalomba , Figs 6 H–K); coronal markings (when present) confined to tylus or forming transverse band between antennal ledges. Head usually angularly produced; pronotum usually sparsely punctate (shiny in T. bimaculatus Synave ); tegminal veins obscure or irregularly patterned with black, sometimes black at tips ( Figs 7 E– Q); pecten of 2nd tarsomere of hind leg with 5–6 black-tipped spines. Slight to distinct sexual dimorphism. Ovipositor 2nd valvulae broad, strongly tapered to pointed tip, without teeth on dorsal margin beyond fused bases ( Figs 11 B–G). Length: 3–5 mm.

Included taxa. There are 10 species in this subgenus, of which half are new to science.

Figures

FIGURE 7. Dorsal aspects of Grellaphia costalis (A), G. robusta sp. nov. (B) and (at slightly larger scale) Tremapterus subgenus Abbalomba (C – D) and T. subgenus Nyanja (E – Q): C, Tremapterus schoutedeni; D, T. facialis; E, T. viduus, female lectotype; F, same, male paralectotype; G, T. perfasciatus sp. nov., male holotype; H, same, female; J, T. virididorsum sp. nov., holotype; K, T. indecisus sp. nov., holotype; L, T. scutellatus sp. nov., holotype; M, T. humilis sp. nov., holotype; N, T. muiri; O, T. bukamensis, lectotype; P, T. johannae, lectotype; Q, T. bimaculatus, lectotype. Scale line: 1 mm.

FIGURE 11. Second valvulae of Taphrotylus (A), Sepullia (M – P) and Tremapterus subgenera Nyanja (B – G), Selenion (H), Tremiziana (J – L), and subgenus Abbalomba (Q – R). A, Taphrotylus insignificans, holotype; B, Tremapterus bimaculatus; C, T. humilis; D, T. perfasciatus; E, T. indecisus; F, T. scutellatus; G, T. virididorsum; H, T. longicaudatus, holotype; J, T. araneosus, paratype; K, T. scapulatus, paratype; L, T. oxythymus, paratype; M, Sepullia fuscolimbata; N, S. viridicollis; O, S. abyssinia; P, S. murrayi; Q, T. typicus; R, T. facialis. Scale line: 1 mm.

FIGURES 3 – 6. Comparison of face (3 – 4 A), tegmen (3 – 5 B), hind wing (3 – 5 C), lateral aspect of head, pronotum and scutellum (6 A – M) and dorsal aspect (N – O) in genera of Clastopterini (3 – 4), Machaerotinae (5), Sepullini (6 A – L, O) and Aphrophorinae (6 M – N): 3 A, Clastoptera atrapicata; 3 B – C, Clastoptera globosa Fowler; 6 A, Clastoptera achatina; 4 B – C, Iba (s. s.) venosa; 5 B – C, Aphrosiphon bauhiniae, from China (1935), 6 B, Iba (Parahindoloides) lumuana, lectotype, 6 C, Beesoniella sylvestris; 6 D, Sepullia fuscolimbatus; 6 E, Tremapterus (Patriziana) somalicus; 6 F, Tremapterus (s. s.) nigropunctatus, lectotype; 6 G, Tremapterus (s. s.) palimpsestus sp. nov.; 6 H, Tremapterus (Abbalomba) schoutedeni, holotype; 6 J, Tremapterus (Abbalomba) facialis, lectotype; 6 K, Tremapterus (Abbalomba) typicus; 6 L, Grellaphia costalis; 6 O, Sepullia viridicollis var. curtula; 6 M – N, Witteella major (M – O from Jacobi 1910, with hind legs of Sepullia corrected to place spines on outer surface).