Lesteva (s.str.) fontinalis Kiesenwetter, 1850

Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, Zootaxa 4966 (4), pp. 401-427 : 408-410

Lesteva (s.str.) fontinalis Kiesenwetter, 1850

( Figs. 9 , 14–15 )

Lesteva fontinalis Kiesenwetter, 1850: 222 ; Ganglbauer 1895: 713, Luze 1903: 195, Portevin 1929: 438, Lohse 1955a: 505, 1960: 3

Lesteva (s.str.) fontinalis: Zanetti 1987: 362

Lesteva (s.str.) fontinalis fontinalis: Tronquet 2006: 33

Lesteva fontinalis gustavi Herman, 2001a: 37 (replacement name of L. fontinalis truncata Lohse, 1960: 5 ) syn.n.

Lesteva nigra Kraatz, 1857: 934

Lesteva cavernicola Scheerpeltz, 1933: 1063

Type material examined. Lectotype (here designated) of Lesteva fontinalis Kiesenwetter, 1850 ♂ [The specimen is glued sideways on small triangular card; head, pronotum and left legs are missing]: ‘Kiesenwetter’ <printed>, ‘fontinalis | M[ons]. Serr[atus Cataloniae].’ <handwritten>, ‘Sammlung | Cl. Müller’ <printed>, ‘ Holotypus | (vermutlich!) [handwritten] | Lesteva [handwritten] | fontinalis Kiesw. [handwritten] | Staatssamml.München’ <red, printed>, ‘zweifellos die Type, | da nur dies eine Ex. | von mon serrat | St..[…illegible…]’ <handwritten>, ‘Zool. Staatsslg. | München’ <blue, printed>, ‘ LECTOTYPE | Lesteva fontinalis | Kiesenwetter, 1850 | Shavrin A.V. des. 2021’ <red, printed> ( ZSM).

Paratype of Lesteva fontinallis truncata Lohse, 1960 ♂ [aedeagus glued under the specimen]: ‘ Espagne: Estremadura | Guadalupe (scierie) | rio Guadalupejo | V—1958, G.Fagel’ <printed>, ‘ Lesteva [handwritten] | fontinalis [handwritten] | ssp truncata [handwritten] | det.Dr.G.A.Lohse’ <printed>, ‘Para- | type’ <red, printed>, ‘Coll. | G. A. Lohse | MHNG-1994’ <printed>, ‘ PARATYPE | Lesteva | fontinalis | truncata ssp. n. | Det Lohse 1960 ’ <yellow, printed>, ‘ PARATYPE | Lesteva fontinalis | gustavi nom.n. | Det. Herman 2001’ <printed>, ‘MHNG | ENTO | 00087335’ <printed> ( MHNG). Paratype ♀, same labels except for ‘MHNG | ENTO | 00087336’ ( MHNG). Paratype ♂: ‘ Espagne: Estremadura | Guadalupe | Fuente del Trincho | V—1958, G.Fagel’ <printed>, ‘fontinalis [handwritten] | ssp. truncata [hadwritten] | det.Dr.G.A.Lohse’ <printed>, ‘Coll. | G. A. Lohse | MHNG- 1994’ <printed>, ‘ Lesteva fontinalis | infrasp. gustavi nom.n. | Det. Herman 2001’ <printed>, ‘MHNG | ENTO | 00087337’ <printed> ( MHNG). All paratypes with additional printed label: ‘ Lesteva (str.) fontinalis Kiesenwetter, 1850 | Shavrin A.V. det. 2021’.

Material examined. SPAIN: ASTURIAS: 2 ♂♂: Puerto Pajares. A. Kricheldorff leg. ( NMPC) ; PAIS VASCO: 1 ♂: Bilbao ( MHNG) ; CATALUñA: 1 ♂: Col d`Ares ( Spain Pyren. ). 31.05.1971. G. Lohse leg. ( MHNG) ; 2 ♂♂: Barcelona, Monserrat. 22.06.2006. L. Fancello leg. (cZan) ; 1 ♂: Sierra de Montseny. 13.05.1981. P. Zwick leg. ( MHNG) ; 2 ♂♂, 2 ♀♀: Seo de Urgel ( Arfa ), de la Coma. 05-06.1962. G. Fagel leg. ( MHNG) ; 1 ♀: Seo de Urgel (graviers du Segre). 05-06.1962. G. Fagel leg. ( MHNG) ; ARAGÓN: 1 ♂: Ardisa ( MHNG) ; 3 ♂♂, 2 ♀♀: Ordesa, Huesca. 31.05.1965. A. Comellini leg. ( MHNG) ; 12 ♂♂, 2 ♀♀: Teruel, Albarracin. 1200 m a.s.l. 23.05.1993. A. Matern (cA, cW, cSh) ; 1 ♂, 1 ♀: Mezquita de Jarque. 18.06.1970. A. Comellini leg. ( MHNG) ; CASTILLA Y LÉON: 5 ♂♂, 5 ♀♀: Sierra de la Demanda. H. Meybohm leg. ( MHNG) ; 1 ♂: Santa Colomba de Somoza, Astorga. 31.05.1970. A. Comellini leg. ( MHNG) ; MADRID: 1 ♂: Escorial ( MHNG) ; CASTILLA LA MANCHA: 3 ♂♂: Las Torcas. 1200 m a.s.l. 23.05.1962. H. Freude leg. ( MHNG, ZSM) ; ANDALUSÍA: 3 ♂♂: Sierra de Cazorla, sources of Guadalquivir. 25.06.1975. M. Daccordi leg. (cZan) ; 1 ♀: Sierra de Cazorla. 23.06.1975. M. Daccordi leg. (cZan) ; 2 ♂♂: same locality. 27.05.2009. T. Struyve leg. (cZan) ; 1 ♀: Sierra de Segura, Guadalquivir , 37°50’N - 2°59’W. 2276 ft. 21.05.2002. Starke leg. (cA) ; 4 ♂♂, 2 ♀♀: Sierra Magina , 37°72’N - 3°46’W. 2000 m a.s.l. 03- 04.05.2014. A.V. Shavrin leg. (cSh) ; 1 ♂, 2 ♀♀: ‘ Kiesenwetter’, ‘ Jaen (Sierra)’, ‘ Sammlung Cl. Müller’, ‘ fontinalis Kiesw’ ( ZSM) ; 1 ♂: ‘ Jaen’ , ‘ Sammlung Cl. Müller’, ‘ fontinalis Kiesw’ ( ZSM) ; 1 ♂: Jaen ( MHNG) ; 1 ♂: Jaen, Jaen-Valdepenas Rd. , 18 km S Jaen. 1000 m a.s.l. 25.05.1967. M.E. Bacchus & D. Levey ( MHNG) ; 1 ♀, 1 ex.: ‘S.d. Jaen’, ‘ Sammlung Cl. Müller, ‘ fontinalis ?’ ( ZSM) ; 1 ♂: Cazorla. 800-2000 m a.s.l. 05.1943. G. Frey & C. Koch ( MHNG) ; 1 ♂: ‘ fontinalis Ksw type. M… [illegible]’, ‘…[illegible] prés Granada’, ‘ Andalousie’ ( IRSN) ; 1 ♂: ‘ Spanien, 1935 Sierra Nevada 2000 m, 16/6 O. Lundblad’, ‘ Lesteva fontinalis Kiesw. Det. Scheepeltz 1964 ’ ( SMNH) ; 1 ♂, 1 ♀: ‘ Süd. Spanien’, ‘ Andalusien’ ( NMPC) ; LOCALITY NOT SPECIFIED: 2 ♂♂: ‘ Spanien Scriba’ , ‘ Coll. K. Neumann’ ( SF) .

Additional material. FRANCE: 1 ♀: ‘St. Jean de Luz Basse-Pyrenees’ ( NMPC); MOROCCO: 1 ♂, 1 ♀: ‘Rif Ketama’ ( MHNH); LOCALITY NOT SPECIFIED: 1 ♂, ‘Pyrén[ees].’, ‘ fontinalis ’, ‘ L. pandellei J. Bohač det. 1980’ ( NMPC); 1 ♂: Pyrenees ( MHNG).

Remarks. Lesteva fontinalis was originally described based on an unspecified number of syntypes from “Mons serratus [Montserrat] Cataloniae”. During the study of Lesteva from ZSM, the first author found a strongly damaged male, which undoubtedly belongs to the type series. Consequently, this specimen is designated as the lectotype. Lohse (1960) described L. fontinalis truncata as “kleine, kurzflügelige und kleinäugige Rasse” from Central Spain. Herman (2001a) replaced the preoccupied name with L. fontinalis gustavi Herman, 2001 . Based on the study of the type and additional material from Spain and other countries, we did not find pronounced morphological differences between them. Body size, coloration, and the proportions of temples and elytra of L. fontinalis are variable. Thus, the new synonymy is proposed here. Habitus as in Fig. 9 . Aedeagus as in Figs. 14–15 . Lesteva fontinalis is known from Switzerland, France, Spain, and North Africa ( Schülke & Smetana 2015). In North Africa it was recorded from Morocco, Algeria, and Tunisia (Fauvel 1871 [=1872], 1878, 1886, 1902, Champion 1898), but all these records require confirmation. It was recorded from Spain by Fauvel (1871a [= 1872]) and Lohse (1955 a, 1960).

Figures

FIGURES 9–13. Habitus of Lesteva: 9–L. fontinalis (Spain: Despenaperros), 10–L. hanseni (Spain: Ronda), 11–L. lusitana (Portugal: Sierra de Estrela), 12–L. mariei (France: Casteil), 13–L. mateui (Spain: Sierra Nevada). Scale bar: 1.0 mm.

FIGURES 14–19. Aedeagus of Lesteva (14–15: L. fontinalis (Spain: Sierra Magina), 16–17: L. hanseni (Spain: Ronda), 18–19: L. lusitana (Portugal: Sierra de Estrela)): 14, 16, 18–parameral view, 15, 17, 19–lateral view. Scale bar: 0.1 mm.