Amynthas papulosus papulosus ( Rosa, 1896 )

Nguyen, Tung T., Nguyen, Anh D., Tran, Binh T. T. & Blakemore, Robert J., 2016, A comprehensive checklist of earthworm species and subspecies from Vietnam (Annelida: Clitellata: Oligochaeta: Almidae, Eudrilidae, Glossoscolecidae, Lumbricidae, Megascolecidae, Moniligastridae, Ocnerodrilidae, Octochaetidae), Zootaxa 4140 (1), pp. 1-92 : 41

Amynthas papulosus papulosus ( Rosa, 1896)

Perichaeta papulosa Rosa, 1896: 525 , Pl. I, Figs 14–16;

Pheretima papulosa— Michaelsen 1900: 291; Gates 1939: 101; Gates 1972: 206; Do 1994: 101; Nguyen 1994: 60; Le 1995a: 38; Pham 1995c: 93; Nakamura 1999: 39; Thai 2000a: 309; Huynh & Nguyen 2004a: 116; Huynh 2005a: 117; Tran & Ngo 2008: 133; Nguyen V.T. & Tran 2008: 185; Tran & Tran 2010: 8; Pham 2010: 63.

Pheretima (Terrata) papulosa— Thai 1983: 123.

Amynthas papulosus— Sims & Easton 1972: 181; Easton, 1981: 56; Blakemore 2002: 180.

Amynthas papulosus papulosus— Sims & Easton 1972: 236; Blakemore 2007a: 76; Blakemore 2008.

Pheretima composita Gates, 1932: 430 , synonymized by Gates (1972).

Type locality. Indonesia (Sumatra: Balighe ).

Type material. Genoa Museum , Italy.

Examined material. 3 C (SORC-V.155.02) fruit tree garden, Hoai Nhon , Binh Dinh, 16/4/1995, coll. Huynh Thi Kim Hoi ; 2 C and 1 A (SORC-V.155.03) Binh Hiep, Binh Son , Quang Ngai, 15/4/1995, coll. Huynh Thi Kim Hoi ; 4 C and 1 A (SORC-V.155.07) cassava garden, Tien Phuoc, Tien Ky , Quang Nam, 28/1/1998, coll. Pham Thi Hong Ha .

Records from Vietnam. Bac Giang (Luc Ngan; Son Dong: Khe Ro) ; Hai Phong ( Cat Ba Isl. ) ; Hanoi (Ba Vi NP) ; Ha Nam ; Nam Dinh ; Ninh Binh ; Nghe An (Ky Son; Que Phong; Quy Chau) ; Quang Binh (Tuyen Hoa; Quang Trach; Bo Trach; Le Thuy) ; Quang Tri (Dong Ha; Gio Linh; Quang Tri); Thua Thien Hue (Huong Dien; Huong Tra; Hue; Huong Thuy; Nam Dong) ; Da Nang (Hoa Vang; Son Tra; Hai Van) ; Quang Nam (Dien Ban; Que Son; Hiep Duc; Tam Ky; Nui Thanh; Tien Phuoc; Phuoc Son) ; Quang Ngai (Binh Son; Tu Nghia) ; Binh Dinh (Hoai Nhon) ; Gia Lai (Pleiku; Buon Luoi) ; Phu Yen ; Dak Lak (Buon Me Thuot; Cu Mgar) ; Khanh Hoa (Ninh Hoa) ; Ninh Thuan (Ninh Son) ; Binh Thuan (Tuy Phong) (Do 1994; Nguyen 1994; Le 1995a; Pham 1995c, 2010; Huynh & Nguyen 2004a; Huynh 2005a; Tran & Ngo 2008; Nguyen V.T. & Tran 2008; Tran & Tran 2010).

Distribution. Cosmopolitan species; also recorded from Myanmar, Thailand, Indonesia (Sumatra), China, Taiwan ( Gates 1972; Easton 1981; Blakemore 2002).

Remarks. The species seems to be widely distributed in Vietnam, except in the southernmost part. Thai (1983) proposed a new subgenus Terrata of the genus Pheretima , and placed the species under the new subgenus.