Soyedina washingtoni ( Claassen, 1923 ),

Grubbs, Scott A. & Baumann, Richard W., 2019, Soyedina Ricker, 1952 (Plecoptera: Nemouridae) in the eastern Nearctic: review of species concepts, proposed morphology-based species groups, and description of a new species from North Caroli, Zootaxa 4658 (2), pp. 223-250: 236

publication ID

https://doi.org/10.11646/zootaxa.4658.2.2

publication LSID

lsid:zoobank.org:pub:A4D55B3B-9BE9-4918-89C6-8049F5895CF2

persistent identifier

http://treatment.plazi.org/id/B66387BE-F376-B365-FF38-14D4FECCF8CA

treatment provided by

Plazi

scientific name

Soyedina washingtoni ( Claassen, 1923 )
status

 

Soyedina washingtoni ( Claassen, 1923) 

Vernal Forestfly

http://lsid.speciesfile.org/urn:lsid: Plecoptera  .speciesfile.org:TaxonName:606 8

( Figs. 43–50View FIGURES 43–50, 80View FIGURE 80)

Nemoura washingtoni Claassen 1923:287  . Holotype ♂ (CUIC), Mt. Washington, (Coös Co.), New Hampshire, USA

Nemoura washingtoniana Needham & Claassen, 1925:217  (misspelling)

Nemoura (Soyedina) washingtoni washingtoni Ricker, 1952:53 

Soyedina washingtoni Illies, 1966:249 

Nemoura (Soyedina) washingtoni Hitchcock, 1974:112 

Distribution. Canada: NB. USA: CT, MD, ME, NY, PA, WV ( DeWalt et al. 2019).

Material examined. Canada, New Brunswick, Restigouche Co., Robinson Brook, Hwy 17, N Oliver, 16 June 1993, R.W. Baumann & B.C. Kondratieff, 1 male ( BYU)  ; Victoria Co., spring-fed stream, Hwy 105, S Medford, 16 June 1993, R.W. Baumann, C.R. Nelson & B.C. Kondratieff, 2 males, 1 female ( BYU)  . USA, Maine, Kennebec Co., Crystal Spring , S Rome, 31 May 1987, R.W. Baumann & G.Z. Jacobi, 4 males, 5 females ( BYU)  ; Waldo Co, seeps, Webb Road, Thorndike , 3 June 1993, D.S. Potter, 3 males, 1 female ( BYU)  . Maryland, Garrett Co., tributary to Swamp Run, Savage River State Forest , 39.57996, -79.05999, 3 April 1996, S.A. Grubbs, 1 male ( WKUC)GoogleMaps  ; Mill Run, Savage River State Forest , 39.54291, -79.07903, 28 April 1996, S.A. Grubbs, 5 males, 3 females ( WKUC)GoogleMaps  ; spring into Elk Lick Run, Savage River State Forest , 39.60557, -79.09583, 26 April 1996, S.A. Grubbs, 4 males, 6 females ( WKUC)GoogleMaps  . Pennsylvania, Cambria Co., Otto Run , 24 April 1994, J. Earle, 1 male ( JEIC)  ; Erie Co., Sixmile Creek, near Wesleyville , 21 April 1979, R.W. Baumann & E.C. Masteller, 1 male ( BYU)  ; tributary to Fourmile Creek , 10 April 1987, E.C. Masteller, 3 males, 3 females ( BYU)  ; same but 29 April 1987, E.C. Masteller, 3 males, 12 females ( WKUC)  ; McKean Co., tributary Kinzua Creek, Rt 321, 17 April 1979, D.H. Funk, 1 male, 2 females ( SWRC)  ; Warren Co., concrete wall, Kinzua Dam , 22 April 1979, R.W. Baumann & O.S. Flint, 1 male ( BYU)  . West Virginia, Pocahontas Co., East Fork Greenbrier River , 38.61778, -79.67674, 26 April 1987, B.C. Kondratieff & R.F. Kirchner, 1 male ( BYU)GoogleMaps  ; Randolph Co., Taylor Run, Alpena Gap Recreation Area , 38.92050, -79.67470, 4 April 1995, S.M. Clark, 3 males, 3 females ( BYU)GoogleMaps  ; Tucker Co., spring, Fernow Experimental Forest , 16 March 1991, M.B. Griffiths, 4 males, 7 females ( BYU)  .

New province and state records. Canada, Prince Edward Island, Kings Co., Naufrage Springs, near Naufrage River , Hugie Joseph Road , 6 June 1998, C.B. Barr, 2 males ( BYU)  ; Valleyfield River, Hwy 326, S Montague, 20 June 1993, R.W. Baumann & B.C. Kondratieff, 1 male, 1 female ( BYU)  . Nova Scotia, Mary Ann Falls, Cape Breton Highlands National Park , 46.72962, -60.36005, 8 June 2009, K. Heard, J. Orlofske, 1 male ( INHS, Insect Collection 516240)GoogleMaps  ; French Lake, Cape Breton Highlands National Park , 46.73057, -60.86532, 10 June 2009, K. Heard, J. Orlofske, 1 male ( INHS, Insect Collection 515784)GoogleMaps  . USA, Massachusetts, Berkshire Co., seep, Wahconah Falls, Wahconah Falls State Park , 27 April 1991, R.W. Baumann & M.F. Whiting, 1 male, 2 females ( BYU)  . New Hampshire, Coös Co., Cutler Brook, Tuckermans Ravine , Mount Washington , 7 July 1974, S.B. Fiance, 2 males, 9 females & 3 nymphs ( BYU)  ; Tuckermans Ravine, Mount Washington , 23 June 1941, T.H. Frison & H.H. Ross, 2 males, 2 females ( INHS)  ; Grafton Co., Hubbard Brook Watershed, West Thornton , 20 May 1975, S.B. Fiance, 5 males, 6 females ( BYU)  . Vermont, Lamoille Co., West Branch Little River, Hwy 108, near Stowe Ski Area , 6 May 1984, G.R. Fiala, 2 males ( BYU)  .

Remarks. Adults were described in Claassen (1923) and subsequently in Needham & Claassen (1925, with same image of the male in lateral view; their plate 38, fig. 14). The male paraproct was also illustrated in Hitchcock (1974, his fig. 143). The holotype male and allotype female (Glen House, NH—also in Coös Co.) were collected from different localities. The only other Soyedina  species distributed that far northward is S. vallicularia  . The combination of the asymmetric dorsal ( Figs. 43–45, 48View FIGURES 43–50) and ventral sclerites ( Figs. 43–44View FIGURES 43–50), plus the inner epiproct member that is flanged and directed laterally with the flange lip approximately same width apically ( Figs. 44–46View FIGURES 43–50), aligns this species closely with S. carolinensis  and S. merritti  . Soyedina washingtoni  is known from the northern Appalachian Mountain region south to West Virginia and westward across Pennsylvania to streams draining into Lake Erie ( Fig. 80View FIGURE 80).

MD

Museum Donaueschingen

BYU

Monte L. Bean Life Science Museum

SWRC

Stroud Water Research Center

INHS

Illinois Natural History Survey

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Plecoptera

Family

Nemouridae

Genus

Soyedina

Loc

Soyedina washingtoni ( Claassen, 1923 )

Grubbs, Scott A. & Baumann, Richard W. 2019
2019
Loc

Nemoura (Soyedina) washingtoni

Hitchcock 1974: 112
1974
Loc

Soyedina washingtoni

Illies 1966: 249
1966
Loc

Nemoura (Soyedina) washingtoni washingtoni

Ricker 1952: 53
1952
Loc

Nemoura washingtoniana

Needham & Claassen 1925: 217
1925
Loc

Nemoura washingtoni

Claassen 1923: 287
1923