Brotheas granulatus Simon, 1877

Ythier, Eric, 2018, A synopsis of the scorpion fauna of French Guiana, with description of four new species, ZooKeys 764, pp. 27-90: 48-50

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Brotheas granulatus Simon, 1877


Brotheas granulatus Simon, 1877  Fig. 24


Simon 1877, Lourenço 1983, Lourenço 1997b, Fet et al. 2000, Lourenço 2002a, Stockmann and Ythier 2010.


Downstream from Saut Pararé on Arataye river, Approuague tributary, one male, MNHNRS6978, J.P. Gasc coll., VIIVIII/1970. Downstream from Saut Pararé on Arataye river, one male and two females, MNHNRS7390, J.P. Gasc coll., IVV/1979. Downstream from Saut Pararé on Arataye river, one male, MNHNRS8279, J.P. Gasc coll., 12/VII/1977. Downstream from Saut Pararé on Arataye river, at the base of Astrocaryum paramaca  , six males and four females, MNHNRS-8508, J.P. Gasc coll., I/1981. Cayenne, one male and one female (types), MNHNRS-0761. Upper Approuague, Culebane river, in forest, one female, MNHNRS3302, De Floch coll., III/1946. Inini  , Institut Pasteur Station, under a wood log, one male, MNHNRS6267, J.P. Gasc coll., 13/VII/1972. Inini  , Institut Pasteur Station, one male, MNHNRS-6274, J.P. Gasc coll., 16/VII/1972. Inini  , Institut Pasteur Station, one male, MNHNRS6275, J.P. Gasc coll., 14/VII/1972. Saül, one male, one female and six immatures, MNHNRS5288, A.S. Balachowsky coll., 27/X/1969. Saut Sabbat, one female, MNHNRS8203, D. Kopp coll., 9/VII/1976. Saut Sabbat, one male, MNHNRS8204, D. Kopp coll., 15/VII/1975. St. Elie (path), ORSTOM Station, one male and one female, MNHNRS7419, J. Lescure coll., 10/VII/1978. St. Jean du Maroni, five males and one female, MNHNRS3303, R. Benoist coll., 1916. Cacao, one female and one immature, deposited in the MHNG, W. Lourenço leg., II/1989. Cacao, one female, deposited in the MHNG, Chippaux coll., X/1983. Cacao, one male, deposited in the MHNG, T. Fretiag coll., XII/1988. Trinité reserve, inselberg, Aya River, five females and six immatures, deposited in the MNHN, C. Courtial coll., XII/2010. Trinité reserve, Aya River, one female and one male, deposited in the MNHN, C. Courtial coll., X/2009. Nouragues reserve, inselberg, UV collected, on a trunk, one female, deposited in the MNHN, C. Courtial coll., 06-09/XII/2013. Rorota, one female and two males, deposited in the EYPC, EY0026, E. Ythier coll., 28/IX/2005. Matoury, one female and 20 immatures, deposited in the EYPC, EY0035, E. Ythier coll., 03-10/XI/2006. Fourgassier, one male, deposited in the EYPC, EY0046, E. Ythier coll., 03-10/XI/2006. Tresor reserve, three males, deposited in the MNHN, C. Courtial coll., VI/2010. Cayenne and Maroni (?), two females, MNHNRS3296, M. Noirot coll. Cayenne and Maroni (?),one male and one female, MNHNRS-3312, E. Abonnenc coll. Cayenne and Maroni (?),one male, MNHNRS3318, M. Noirot coll. Guyane, one male, MNHNRS-0743, F. Geay coll., 1900. Guyane, four females, MNHNRS3313, M. Tartaire coll., 1921. Guyane, two females, MNHNRS0749.


Species of moderate to large size when compared with the average size of the other species within the genus, ranging from 58 to 68 mm in total length. General coloration brown to blackish brown. Carapace dark brown with several scattered black spots; ocular tubercle dark, almost black. Tergites of same color as the carapace; several paler confluent areas. Sternites brownish yellow, the sternite VII being darker; pectines and genital operculum dark yellowish. Metasomal segments of same color as the tergites; some blackish pigmentation over the carinae. Vesicle reddish yellow with basis of aculeus reddish and tip of aculeus reddish black. Chelicerae dark yellowish with reticular brown spots; basis of fingers and fingers dark browns. Pedipalps blackish brown; chela reddish with longitudinal blackish spots; fingers blackish with tip reddish. Legs yellowish brown with several diffuse dark spots. Carapace weakly emarginated, with strong granulation; ocular tubercle punctate. Tergites with strong granulation similar to the carapace, the tergites VI and VII with stronger granulation. Pectinal tooth count ranging from eight to eleven in male and seven to nine in female. Sternites punctate; spiracles linear, elongated. All metasomal segments with dorsal, latero-dorsal and latero-ventral carinae well-marked; ventral carinae absent on segment I and well-marked on other segments, with spinoid granules on segments IV and V; other cari nae incomplete; intercarinal tegument moderately punctate. Vesicle with four vestigial carinae on ventral side; lateral and ventral sides weakly granulated; aculeus long compared to vesicle, wide at its basis. Pedipalp femur with five well-marked carinae; patella with four carinae; chela with vestigial carinae; dorsal and internal sides of femur well granulated; patella and chela moderately granulated except on internal side of chela where there are stronger granules; dentate margins on movable fingers with six slightly oblique rows of granules, separated by stronger accessory granules.