Riama, GRAY, 2005

Doan, Tiffany M. & Castoe, Todd A., 2005, Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus, Zoological Journal of the Linnean Society 143 (3), pp. 405-416 : 413

KEY TO THE SPECIES OF RIAMA GRAY

1a. Limbs overlapping when adpressed against body.....................................................2

1b. Limbs not overlapping when adpressed............................................................10

2a. Hexagonal dorsal scales.........................................................................3

2b. Quadrangular dorsal scales.......................................................................4

3a. Dorsal scales obtusely hexagonal, preanal pores in males 2................................... R. achlyens

3b. Dorsal scales acutely hexagonal, preanal pores in males 4–6................................... R. shrevei

4a Keeled, striated, or rugose dorsal scales............................................................5

4b Smooth dorsal scales..................................................................... R. laevis

5a. Lateral scale rows 0–6...........................................................................6

5b. Lateral scale rows 7–12..........................................................................9

6a. Transverse rows of dorsal scales 25–35.............................................................7

6b. Transverse rows of dorsal scales 39–43................................................... R. balneator

7a. Lateral scale rows 2–6...........................................................................8

7b. Lateral scale rows 0–1.................................................................... R. striata

8a. Rugose dorsal scales...................................................................... R. vieta

8b. Keeled or striated dorsal scales......................................................... R. hyposticta

9a. Femoral pores on one leg 0–6............................................................. R. oculata

9b. Femoral pores on one leg 9–14........................................................... R. luctuosa

10a. Venter immaculate except near lateral surfaces.....................................................11

10a. Ventral scales with dark pigmentation............................................................12

11a. Preanal pores present in both sexes........................................................ R. inanis

11b. Preanal pores absent.................................................................. R. vespertina

12a. Keeled, striated, or rugose dorsal scales...........................................................13

12b. Smooth dorsal scales.................................................................. R. meleagris

13a. Keeled or striated dorsal scales, dorsal stripes in some individuals, preanal pores present or absent, three or four supraoculars.......................................................14

13b. Keeled dorsal scales, no dorsal stripes, preanal pores present in males and females, four supraoculars............................................................ R. labionis

14a. Transverse rows of dorsal scales greater than 31...................................................15

14b. Transverse rows of dorsals scales 27–31................................................... R. petrorum

15a. Three or four supraoculars, lateral ocelli present or absent, lip barring variable..........................17

15b. Three supraoculars, no lateral ocelli, no barring on lip...............................................16

16a. Supralabial-subocular fusion absent........................................................ R. raneyi

16b. Supralabial-subocular fusion present................................................... R. laudahnae

17a. Superciliary series complete or incomplete, transverse rows of ventrals 19–27, anterodorsal thigh scales smooth, rugose, or striate.................................................18

17b. Superciliary series incomplete, transverse rows of ventral scales 18–20, anterodorsal thigh scales smooth.................................................... R. colomaromani

18a. Keeled or striated dorsal scales, longitudinal rows of dorsal scales 20–34, preanal pores present or absent, femoral pores on one leg of males 0–11................................19

18b. Keeled dorsal scales, longitudinal rows of dorsal scales 21–22, preanal pores absent, femoral pores of males on one leg 3–5........................................... R. columbiana

19a. Keeled or striate dorsal scales, subdigital lamellae on fourth toe 10–18, superciliary series complete or incomplete.........................................................20

19b. Keeled dorsal scales, subdigital lamellae on fourth toe 15–18, superciliary series complete........................................................................... R. orcesi

20a. Transverse rows of ventral scales 19–27, hemipenis with or without small or large spines.................21

20b. Transverse rows of ventral scales 19–20, hemipenis with small spines.......................... R. simotera

21a. Femoral pores of males 0–10, preanal pores of males 0–4, femoral pores of females 0–9, preanal pores of females 0–4....................................................................22

21b. Femoral pores of males 8–9, preanal pores of males 2–4, females lack femoral and preanal pores....................................................................... R. stigmatoral

22a. Hemipenis with small or absent spines, superciliary series usually incomplete...........................23

22b. Hemipenis with large spines, superciliary series complete................................... R. anatoloros

23a. Preanal pores present in both sexes...................................................... R. unicolor

23b. Preanal pores absent................................................................. R. cashcaensis