Goeldia portenha, Almeida-Silva & Brescovit, 2024

Almeida-Silva, Lina M. & Brescovit, Antonio D., 2024, Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae), Zootaxa 5428 (2), pp. 151-193 : 183-185

Goeldia portenha sp. nov.

( Figs 1E–F ; 20 ; 26 )

Type material. Holotype ♂ from Andalgalá (27°36’S 66°19’W), Catamarca, Argentina, 31.I.2986, P. Goloboff coll. ( MACN 16126 ) . Paratypes: 2♀ with same data as holotype ( MACN 16126 ) ; 1♂ from Potrerillos (32°57’S 69°10’59”W), Mendoza, Argentina, 24.III.1979, A. Roig coll. ( IBSP165958 ) ; 3♀ from Tafi del Valle (26°52’S 65°41’W), Tucumán, Argentina, 16.I.1981, A. Roig coll. ( IBSP165949 ) .

Additional Material Examined. BRAZIL. Brasília : Palácio da Alvorada (15°47’33”S 47°49’19”W), 2♂, 1962, R. Schiapelli, & J. M. Viana coll. ( MACN 16071 ) ; ARGENTINA. Tucumán: El Mollar, Reserva Provincial de La Angostura (26°55’S 65°42’W), 2♀, 1 juv. 15.XII.1994, C. Grismado coll. ( MACN 16037 ) ; Tafi del Valle (26°52’S 65°41’W), 19♀, 16.I.1981, A. Roig, coll. ( MACN 16205 ) ; El Infiernillo (26°44’24”S 65°46’16”W), 3♀, 1 juv., 21.IV.2000, P. Goloboff & L. Lopardo coll. ( MACN 16210 ) . La Rioja: La Rioja (29°24’47”S 66°51’21”W), 1♀, XI.1943, J. M. Viana coll. ( MACN16088 ) ; Guayapa (30°6’20”S 66°58’36”W), 1♀, 1 juv., X.1965, E. Maury coll. ( MACN 16082 ) ; Guayapa (30°6’20”S 66°58’36”W), 1♂, X.1963, M. E. Galiano coll. ( MACN 16072 ) ; Patquia (30°03’00”S 66°52’59”W), 4♀ 16–23.X.1963, M. E. Galiano coll. ( MACN 16078 ) ; Chilecito (29°10’S 67°30’W), 1♀, 1955, M. E. Galiano coll., ( MACN 16096 ) ; Cuesta de Miranda (29°20’S 67°48’W), 2♀, VIII.1994, M. Ramírez coll., ( MACN 16048 ) . Catamarca: Mutquin (28°19’S 66°10’W), elev. 200 m, 2♀, 1966, O. de Ferraris coll. ( MACN 16029 ) . La Pampa: (36°37’S 64°17’W) 5♀, 1♂, 20.XI.1934, Pereya coll. ( MACN 16097 ) ; General Duval (38°43’S 66°24’W), 2♀, 13.XI.1974, E. Maury coll. ( MACN 16116 ) ; Parque Nacional Lihuel Calel (37°57’S 65°39’W), 5♀, I. XI.1975, A. Toth & E. Maury coll. ( MACN 16091 ) ; Parque Nacional Lihuel Calel (37°57’S 65°39’W), 2♀ 2 juv., 22.I.1985, M. Ramírez coll. ( MACN 16042 ) ; Parque Nacional Lihuel Calel (37°57’S 65°39’W), 8♀, 1 juv., E. Maury coll. ( MACN 16101 ) ; Santa Rosa (36°37’14”S 64°18’27”W), 1♀, no further data ( MACN 16094 ) ; Parque Nacional Lihuel Calel, cerro Fortaleza , (38º1’14”S 65º35’32”W), elev. 419 m, 1♂, 3♀, 5 juv., 11.X.2014, A.D. Brescovit, A.J. Santos, L.N. Piacentini, I.L.F. Magalhães ( MACN 34592 ) ; Parque Nacional Lihuel Calel, cerro de la Sociedad Científica Argentina (38º0’25”S 65º35’46”W), elev. 348 m, 1♀, 11.X.2014, A.D. Brescovit, A.J. Santos, L.N. Piacentini, I.L.F. Magalhães ( MACN 34547 , Fig. 1F ) . Salta: between Santa Maria (22°08’S 62°49’W) y Cafayate (26°05’S 65°58’W), 1♂, 19.X.1948, W. Partidge coll. ( MACN 16134 ) ; Cerro Chuscha (26°08’56”S 66°13’51”W), 6 km NW El Cafayate, 3♀, 17.VII.1995 M. Ramírez & P. Goloboff coll. ( MACN 16036 ) ; La Salamanca , 7 Km S Alemanía (25°37’27”S 65°36’47”W), 18.VII.1995, P. Goloboff & M. Ramírez coll. ( MACN 16069 ) ; Neuquén: Neuquén (38°58’S 68°4’W), 1♂, IX.1973, O. de Ferraris coll. ( MACN 16100 ) ; San Juan: Las Tumanas (30°51’53”S 67°19’20”W), 20 Km S Valle Fertil, 1♀, 1 juv., 14.IV.1979, A. Roig coll. ( MACN 16129 ) ; Mendoza: Potrerillos (32°57’S 69°10’59”W), 1♂, 1♀, 24.III.1979, A. Roig coll. ( MACN 16206 ) ; Papagayos (32°49’19”S 68°53’47”W), 2♀, 3 juv., IX.1981, A. Gonzalos coll. ( MACN 16035 ) .

Etymology. The specific epithet is a Brazilian Portuguese noun in apposition for the people from Buenos Aires, capital of Argentina.

Diagnosis. Goeldia portenha differs from other species of the genus by the presence of a pair of dorsal white spots (guanine deposits) on the opisthosoma, which are very strong in males but faint or absent in females ( Fig. 1E–F ). Males differ from other species by the tegular process usually with a subapical constriction ( Fig. 20A–B ). The female can be diagnosed by the epigynum with the rim almost vertical and fertilization ducts far from each other twice its length ( Fig. 20E ).

Description (Male, MACN 16126). Coloration: carapace brown, cephalic area slightly darker than thoracic area; sternum, endites and labium brown; chelicerae reddish brown. Legs: femur I as cephalic area and other leg segments yellowish brown. Opisthosoma brown with a dorsal pair of guanine deposits spots ( Fig. 1E ). Total length 5.25. Carapace: length 2.60; width 1.55; height 1.00. Clypeus: height 0.15. Opisthosoma: length 2.65; width 3.1. Eye diameters and interdistances:AME 0.08; ALE 0.10; PME 0.09; PLE 0.1; AME–AME 0.14; AME–ALE 0.09; PME– PME 0.09; PME–PLE 0.19; AME–PME 0.09; ALE–PLE 0.05. Chelicerae paturon: length 1.16. Leg measurements: I: femur 2.22/ patella 0.81/ tibia 2.00/ metatarsus 1.88/ tarsus 0.91/ total 6.47; II: 1.94/ 0.78/ 1.41/ 1.63/ 0.72/ 5.03; III: 1.44/ 0.66/ 1.13/ 1.22/ 0.59/ 5.03; IV: 1.88/ 1.75/ 0.78/ 1.63/ 0.63/ 6.66. Leg formula 1243. Leg spination: femur I p1ap; tibia I v0-2-0-2, r1-1-0-0-0; II p1ap; III v2sap, p1ap, r1ap; IV v0-0-1p-2, p1ap, r1ap; metatarsus I v1r-0- 1r- 0-1m, p1ap, r1ap; II v1r-0-1-1m- 1m, p1ap, r1ap; III v1r-0-2- 1m, p1ap, r1ap, d2ap; IV v2-0-1r-2-1p- 1m, p1ap, r1ap, d1ap. MLT apex partially sclerotized ( Fig. 20C ). Patelar apophysis conic and slightly curved posteriorly. Pars pendula thickening subdistal ( Fig. 20C ).

Female (MACN 16126). Coloration: as in male except for the opisthosoma dorsal spots, which are beige and not white guanine deposits ( Fig. 1F ). Total length 5.00. Carapace: length 2.65; width 1.75; height 1.0. Clypeus: height 1.6. Opisthosoma: length 2.55; width 2.05. Eye diameters and interdistances: AME 0.08; ALE 0.13; PME 0.09; PLE 0.11; AME–AME 0.10; AME–ALE 0.11; PME–PME 0.14; PME–PLE 0.21; AME–PME 0.13; ALE–PLE 0.08. Chelicerae paturon: length 1.14. Leg measurements: I: femur 1.88/ patella 0.92/ tibia 2.84/ metatarsus 1.4/ tarsus 0.54/ total 7.58; II: 1.72/ 0.8/ 1.56/ 1.36/ 0.64/ 6.08; III: 1.44/ 0.8/ 1.36/ 1.56/ 0.64/ 5.80; IV: 1.78/ 1.59/ 0.91/ 1.22/ 0.56/ 6.06. Leg formula 1243. Leg spination: femur I p1ap; tibia III r1ap; metatarsus I v1map, II–III v1map, p1ap, r1ap; IV v0-1p-1map, p1ap, r1ap. Copulatory ducts almost invisible at the copulatory opening area ( Fig. 20E ). Copulatory ducts short with apex dorsally bent ( Fig. 20F ). Vertical spermathecae taller than copulatory ducts and L-shaped ( Fig. 20F ). Horizontal spermathecae straight ( Fig. 20F ).

Variation. Length (8♂): total 4.95–5.48; carapace 2.47–2.69; femur I 1.97–2.34. (10♀): total 4.85–6.08; carapace 2.38–3.10; femur I 1.76–2.05. Some females lack the white spots on the dorsum of the opisthosoma. Tegular process sometimes more curved as represented in Fig. 20A and patellar apophysis proximally more sinuous ( Fig. 20C ).

Distribution. Known from Brasília , Brazil, and Argentina ( Fig. 26 ).

Figures

FIGURE 1. A–F. Goeldia spp. habitus:A. Goeldia mirim sp. nov., female, from state of Rondônia, Brazil; B–D. Goeldia nigra Mello-Leitão from São Paulo, Brazil: B. ♀; C. ♀, with eggsac and cribellate silk; D. ♂; E–F. Goeldia portenha sp. nov. from La Pampa, Argentina: E. ♂; F. ♀.

FIGURE 20. A–F. Goeldia portenha sp. nov. A–D. Male left palp; E–F Female Epigynum: A. Tegular process three views. B. Palp ventral view; C. Tibia and patella, dorsal view; E. Epigynum ventral view; F. Vulva, dorsal view.

FIGURE 26. Distribution of Goeldia spp. in southern South America: Goeldia leechi (square—blue); Goeldia portenha (star— red); Goeldia nigra (circle—orange) and Goeldia bagumbubu (triangle—yellow). Type species are marked as black versions of each symbol when there are records to multiple localities.