18.
Pontodrilus litoralis (Grube, 1855)
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Lumbricus litoralis Grube, 1855: 127
;?[
Pontoscolex arenicola Schmarda, 1861: 11
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(?part);
Pontodrilus marionis Perrier, 1874: 1582
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;
Pontodrilus bermudensis Beddard, 1891: 96
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;
Cryptodrilus insularis Rosa, 1891: 387
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, fig. 11 (of left prostate);
Pontodrilus arenae Michaelsen, 1892: 222
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, fig. 9;
Pontodrilus sp.
Michaelsen, 1894: 184;
Pontodrilus hesperidum Beddard, 1894: 37
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;
Pontodrilus insularis
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: Beddard, 1895 a: 471;
Pontodrilus michaelseni Eisen, 1895: 73
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;
Pontodrilus ephippiger Rosa, 1898: 277
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, 281;
Pontodrilus matsushimensis Iizuka, 1898: 21
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, Pl. II;
Pontodrilus ephippiger laysanianus Michaelsen, 1899: 217
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;
Pontodrilus matsushimensis chathamianus Michaelsen, 1899: 220
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;
Pontodrilus michaelseni hortensis Eisen, 1900: 129
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, 241, figs.;
Pontodrilus laccadivensis Beddard, 1903: 374
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;
Pontodrilus crosslandi Beddard, 1906: 561
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;
Pontodrilus albanyensis Michaelsen, 1907
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a: 185;
Pontodrilus bermudensis ephippiger
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: Stephenson, 1915: 145;
Plutellus (Pontodrilus) bermudensis
: Michaelsen, 1922: 22; 1935: 106;
Pontodrilus albanyensis var. cygni Jackson, 1931: 94
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;
Plutellus (Pontodrilus) matsushimensis indica Michaelsen, 1935: 106
;
Pontodrilus gracilis Gates, 1943: 100
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.
Pontodrilus litoralis
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: Beddard, 1895 a: 469 (syn.
marionis
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); Michaelsen, 1900: 180 (syn.
marionis
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); Cognetti, 1901: 404; Michaelsen, 1910 c: 89 (syn.
marionis
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,
crosslandi
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,
albanyensis
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); Easton, 1984: 114, fig. 1f [syn.
arenicola
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(part.),
marionis
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,
bermudensis
,
insularis
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,
arenae
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,
hesperidum
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,
michaelseni
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,
ephippiger
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,
bermudensis
ephippiger
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.- Stephenson, 1915,
matsushimensis
,
ephippiger
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laysanianus (syn.
michaelseni
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hortensis),
matsushimensis
chathamianus,
michaelseni
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hortensis, P. sp. Michaelsen, 1894, P. (P.) sp. Michaelsen, 1930,
laccadivensis
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,
crosslandi
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,
albanyensis
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, P. sp. Michaelsen, 1913,
albanyensis
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cygni,
matsushimensis
indica
, P. sp. Gates, 1936,
gracilis
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].
Distribution: CHI: Galapagos Islands ( Blakemore, 2002). COL: Cartagena ( Michaelsen, 1910). VEN: Morrocoy National Park: Playa Mero; Cayo Cico; Cayo Sombrero ( Easton, 1984; Righi, 1989; Zicsi, 1995a). BRA: RS ( Michaelsen, 1900b, 1910); SC: Florianópolis; São Francisco Island; SP ( Lüderwaldt, 1927; Righi, 1967b; Brown and James, 2006): Santos ( Michaelsen, 1910); Itanhaém ( Lüderwaldt, 1927); Cananéia, in mangrove ( Righi, 1968); RJ ( Moreira, 1903); Angra dos Reis, in mangrove ( Righi, 1968); PE: Itamaracá Island (Brown et al., 2006) ( Figure 3D
View Figure 3
).
Further distribution: Atlantic, Indian, and Pacific oceans, fully marine ( Easton 1984).
Remarks: Blakemore et al. (2012) added this species to Korean species biodiversity list as an exotic species from Korea.