Kachinoperla zwicki, Chen, 2022

Chen, Zhi-Teng, 2022, The larval morphology of a new mid-Cretaceous stonefly and its systematic position in Plecoptera, Arthropod Systematics & amp; Phylogeny 80, pp. 423-438 : 423

publication ID

https://dx.doi.org/10.3897/asp.80.e82549

publication LSID

lsid:zoobank.org:pub:1B107F79-F4F7-4342-946E-444777653E0A

persistent identifier

https://treatment.plazi.org/id/924CBD2C-1BAF-4410-AF89-4C81AD28D94A

taxon LSID

lsid:zoobank.org:act:924CBD2C-1BAF-4410-AF89-4C81AD28D94A

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Kachinoperla zwicki
status

sp. nov.

Kachinoperla zwicki sp. nov.

Figs 1 View Figure 1 , 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5

Etymology.

The species is named after Dr. Peter Zwick, who has made substantial contributions to the knowledge of Plecoptera .

Type material.

Holotype: exuvia (No. CZT-PLE-MA13), deposited in the Insect Collection of Jiangsu Uni-versity of Science and Technology (ICJUST).

Locality and horizon.

Amber mine near Noije Bum Village (26°20′N, 96°36′E), Tanai Township, Myitkyina District, Kachin State, northern Myanmar. Horizon unnamed, mid-Cretaceous, Upper Albian to Lower Cenomanian (98.79 ± 0.62 Ma).

Diagnosis.

As for the genus.

Description.

Body robust and slightly oblate (Fig. 1 View Figure 1 ). Surface generally brown to dark brown, with hairs and setae of various size and shape (Fig. 1 View Figure 1 ). - Head: Head wider than long (Fig. 2 View Figure 2 ), anterior area of ecdysial suture dark brown. Anterior ocellus tiny; compound eyes large and rounded, length approximately 0.5 × head length; postocular area fringed with moderately long setae. Occiput pale brown (Fig. 2A View Figure 2 ), with a transverse row of short spinules, the irregular spinules scattered and sinuous. Antennae partially preserved, dark brown and slender, each antennal segment wider than long, near glabrous, apically fringed with sparse short spines. Mouthparts of predaceous type (Fig. 2 View Figure 2 ). Glossae of labium approximately as long as paraglossae. Maxilla well-developed; cardo broad, lateral surface covered with dense short spines; stipes elongated, with rounded, glabrous outer margin, boundary with lacinia invisible; lacinia long, outer margin circular, with two giant apical teeth and an inner comb of moderate marginal spines basal to the second apical tooth; the second shorter apical tooth of left lacinia truncate. Maxillary palp with four visible elongated palpomeres, distal palpomere much thinner and approximately half in length of previous segment. Mandible with three pointed apical teeth, anterior half of inner margin with a marginal comb of cylindrical, stout spines posterior to the base of apical teeth; molar area unmodified. Labial palp similar in length to maxillary palps, with three elongated palpomeres; distal palpomere thinner and slightly shorter than previous segments, apex obliquely truncate. All visible maxillary and labial palpomeres glabrous. - Prothorax: Pronotum wide and short with obtuse corners (Fig. 2A View Figure 2 ), mostly dark brown, margins darker, anterior margin longer than posterior margin; surface glabrous, posteromedial area near the median suture with two irregular pale spots, posterior margin fringed with long cylindrical spines (Fig. 2A, B View Figure 2 ). - Meso- and metathorax: Wing pads with anterior and posterior parts pale brown, medial part dark brown (Fig. 3 View Figure 3 ); all wing pads with same size on meso- and metathorax; each wing pad with a transverse row of moderately long spines near anterior margin, dorsal surface and lateral margin mostly covered with dense patch of collapsed short spinules, apical one third of wing pad near glabrous; posterior notal contour not apparent. Thoracic gills not visible (Fig. 3D View Figure 3 ). - Legs: Legs covered by irregular patches of setae with various size and shape (Fig. 4 View Figure 4 ). Coxae and trochanters dorsally with moderate spines along lateral margins. Femora dorsally and ventrally covered with long cylindrical spines along anterior margin and posterior areas, posterior margin with dense long swimming hairs. Tibiae with several moderate spines and dense long swimming hairs along posterior margin, dorsal and ventral surfaces without obvious spines. Two giant and several smaller tibial spurs present. Tarsi with first two tarsomeres very short, third one approximately 2 × longer than first two combined and apically with several spines. Tarsal segments with sparse long swimming hairs along outer margin. Claws sharp and glabrous, inner margin with pronounced denticles. - Abdomen: Abdominal segments dark brown, each segment covered and posteriorly fringed with long cylindrical spines (Fig. 5 View Figure 5 ). Two distinct abdominal gill remnants present anterior to the first segment (Figs 3D View Figure 3 ; 5C View Figure 5 ). Tergum 10 strongly projected backwards, near triangular in shape, exceeding posterior margin of paraprocts (Fig. 5B View Figure 5 ). Paraprocts subtriangular, inner margins straight and with dense giant spines, posterior margins rounded and glabrous, posterior margins blunt (Fig. 5D View Figure 5 ). Cerci partially preserved, dark, inner and outer margins fringed with sparse rows of giant spines, each segment slightly longer than wide (Fig. 5B View Figure 5 ).

Measurements.

BL 9.5 mm, HL 2.0 mm, HW 2.5 mm, ATL 4.0 mm (partially preserved), PL 1.2 mm, PW 2.0 mm, WL 3.0 mm, FLL 5.0 mm, MLL 6.0 mm, HLL 7.0 mm, ABL 5.0 mm, ABW 1.5 mm, CL 2.0 mm (partially preserved).