Euwallacea gravelyi (Wichmann, 1914) stat. res.

Smith, Sarah M., Beaver, Roger A. & Cognato, Anthony I., 2020, A monograph of the Xyleborini (Coleoptera, Curculionidae, Scolytinae) of the Indochinese Peninsula (except Malaysia) and China, ZooKeys 983, pp. 1-442 : 1

Euwallacea gravelyi (Wichmann, 1914) stat. res. Fig. 55E, F, K

Xyleborus gravelyi Wichmann, 1914: 411.

Euwallacea gravelyi (Wichmann): Saha and Maiti 1996: 815.

Xyleborus ovalicollis Eggers, 1930: 193. Synonymy: Saha and Maiti 1996: 815.

Xyleborus barbatomorphus Schedl, 1951a: 72. syn. nov.

Type material.

Holotype Xyleborus barbatomorphus (NHMW), paratype (NHMW, 1). Holotype Xyleborus ovalicollis (FRI).

New records.

Bhutan: W. Paro distr., Gedu, 2100 m, 17-26.vi.1988, C. Holzschuh (RABC, 1). China: Yunnan, Lincang, Genma, 12.xii.2018, Y. Li, ex rubber tree (MSUC, 1). India: Arunachal Pradesh, 0.3 km SSE of Dirang, 27°20'32"N, 92°16'17"E, 1550 m, 27.iv-1.v.2008, H. Podskalská & P. Šipek (NHMP, 1); as previous except: Etalin vicinity, 28°36'56"N, 95°53'21"E, 700 m, L. Dembický, 12-25.v.2012 (ZFMK, 2). Meghalaya, Nokrek N.P., 3 km S Darbokgiri, 25°27'N, 90°19'E, 1400 m, 26.iv.1999, Dombický, Pacholátko (RABC, 1). Laos: Bolikhamzai, Ban Nape (8 km NE), 18°21'N, 105°08'E, 600 m, 1-18.v.2001, V. Kuban (NHMB, 6; RABC, 3). Champasak, Bolavens Plateau, waterfall ~ 2 km E Tao Katamtok, 15°08.1'N, 106°38.8'E, 415 m, 10-12.v.2010, J. Hájek (NHMP, 1). Taiwan: Nantou, Sun Moon Lake, C.-S. Lin 15.v.2014 (MSUC, 1). Thailand: Chiang Mai, Doi Pui, 1400 m, 6-10.vi.2005, W. Puranasakul (RABC, 1). Vietnam: Tonkin, Hoa-Binh, 1940, A. De Cooman (MNHN, 1).

Diagnosis.

4.6-5.7 mm long (mean = 5.21 mm; n = 8); 2.6-2.75 × as long as wide. This species is distinguished by its large size and elongate form; protibiae less distinctly triangular than E. destruens , and with 7-9 denticles in apical 1/2; declivity commencing at approximately midpoint, evenly curved from disc into declivity; and declivity usually appearing flat in lateral view.

Similar species.

Euwallacea destruens .

Distribution.

Bhutan*, China* (Yunnan), India (Arunachal Pradesh*, Assam, Meghalaya*, West Bengal), Laos*, Myanmar, Taiwan, Thailand*, Vietnam*.

Host plants.

Polyphagous ( Saha and Maiti 1996).

Remarks.

This species was included in Xyleborus by Wood and Bright (1992) but was transferred to Euwallacea by Saha and Maiti (1996) with Xyleborus ovalicollis as a synonym. The location of the two syntypes of E. gravelyi is not known, but Wichmann’s description is sufficiently detailed for us to be able to confirm the synonymy given by Saha and Maiti (1996). Maiti and Saha (2004) included both species as synonyms of Euwallacea wallacei (Blandford), presumably following the placement of E. ovalicollis as a synonym of E. wallacei by Schedl (1970a) and Wood (1989). However, E. wallacei is a distinct species only superficially similar to E. gravelyi . In E. gravelyi , the eyes are of normal size, and the upper part is smaller than the lower; in E. wallacei , the eyes are unusually large and extend onto the frons, the upper and lower parts are of equal size. In E. gravelyi , the protibiae bears 7-9 small denticles in the apical 1/2; in E. wallacei , there are only five large denticles.

Xyleborus barbatomorphus was given as a synonym of E. wallacei by Beaver et al. (2014), but is in fact conspecific with E. gravelyi and is here placed in synonymy. E. wallacei is not known to be present in the area covered by this study.

Figures

Figure 55. Dorsal, lateral and declivital view of Euwallacea funereus, 3.45 - 3.7 mm (A, B, I), E. geminus holotype, 2.7 - 2.8 mm (C, D, J), E. gravelyi, 4.6 - 5.7 mm (E, F, K), and E. interjectus, 3.5 - 3.9 mm (G, H, L).