Pholcus barisan Huber

Bernard A. Huber, Joseph K. H. Koh, Amir-Ridhwan M. Ghazali, Kamil A. Braima, Olga M. Nuñeza, Charles Leh Moi Ung & Booppa Petch, 2016, New leaf- and litter-dwelling species of the genus Pholcus from Southeast Asia (Araneae, Pholcidae), European Journal of Taxonomy 200, pp. 1-45: 21-25

publication ID 10.5852/ejt.2016.200

publication LSID

persistent identifier

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scientific name

Pholcus barisan Huber

sp. nov.

Pholcus barisan Huber  sp. nov.

Figs 63–66View Figs 59 – 66, 72–76View Figs 72 – 76, 80View Figs 77 – 80, 90–92


Easily distinguished from most similar known relatives (representatives of the P. ethagala  group on the Malay Peninsula) by morphology of male palps ( Figs 72–73View Figs 72 – 76; very long dorsal process of procursus; relative straight male palpal trochanter apophysis similar to P. uludong Huber  sp. nov. and P. phui  but shorter; shape of relatively small appendix), by modified male clypeus (pair of small dark processes; Fig. 80View Figs 77 – 80), by male eye stalks directed more towards lateral ( Fig. 80View Figs 77 – 80), and by details of female internal genitalia ( Figs 75, 76View Figs 72 – 76; anterior undulating sclerite; large posterior semicircular structure).


The species name is derived from the type locality; noun in apposition.

Material examined


INDONESIA: ♂, ZFMK ( Ar 15708), Sumatra, Lampung, Bukit Barisan (5.528°S, 104.424°E), 550– 600 m a.s.l., forest near road, leaf litter, 24 Oct. 2009 (S. Sutono)GoogleMaps  .

Other material

INDONESIA: 6 ♀♀, 3 juvs, ZFMK ( Ar 15709), same data as holotypeGoogleMaps  ; 4 ♀♀, 1 juv., in absolute ethanol, ZFMK ( Ind 153), same dataGoogleMaps  .


Male (holotype)

MEASUREMENTS. Total body length 3.3, carapace width 0.9. Leg 1: 33.2 (7.6 + 0.4 + 7.7 + 15.6 + 1.9), tibia 2: 4.5, tibia 3: 2.6, tibia 4: 4.0; tibia 1 L/d: 96. Distance PME-PME 510 µm, diameter PME 100 µm, distance PME-ALE ~30 µm; AME absent.

COLOR. Carapace pale ochre-yellow with large dark brown mark posteriorly, dark median line, ocular area, stalks and clypeus also dark brown; irregular black marks in place of AME; sternum whitish; legs ochre-yellow with darker brown to black patellae and tibia-metatarsus joints; abdomen ochre-gray with some darker marks dorsally and laterally.

BODY. Habitus as in Figs 63–64View Figs 59 – 66; ocular area raised, each triad on long stalk directed laterad and dorsad ( Fig. 80View Figs 77 – 80); carapace without median furrow; clypeus with pair of small dark processes at rim; sternum wider than long (0.60/0.53), unmodified.

CHELICERAE. As in Fig. 74View Figs 72 – 76, with single pair of frontal proximal apophyses; without distal modification; without stridulatory ridges.

PALPS. As in Figs 72–73View Figs 72 – 76; coxa with indistinct ventral hump; trochanter with very long almost straight retrolatero-ventral apophysis and small retrolateral process; femur with indistinct retrolateral hump; tarsus without dorsal elongation; procursus complex, proximal part with distinctive dorsal process forming retrolateral groove distally, ventral side heavily sclerotized, dorsal side semitransparent; hinged distal part with large prolatero-dorsal flap and weakly sclerotized prolateral process lying parallel to procursus; bulb with rather simple appendix, long slender embolus, without uncus.

LEGS. Without spines and curved hairs; few vertical hairs; retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium absent on tibia 1, present on other tibiae; tarsus 1 pseudosegments very indistinct, apparently irregular.


In general similar to male ( Figs 65–66View Figs 59 – 66) but eye triads not on stalks and much closer together than in male (PME-PME distance: 240 µm); dark posterior mark on carapace smaller than in male, without median dark line on carapace; with pair of small black marks in area of AME (but without lenses); clypeus unmodified and only in upper third dark brown. Tibia 1 in 6 females: 6.0–6.9 (mean 6.4). Some females (and juveniles) with laterally strongly widened abdomen ( Fig. 65View Figs 59 – 66). Epigynum weakly sclerotized (Fig. 90), with small ‘knob’ at posterior rim ( Fig 75View Figs 72 – 76); internal anterior arc undulating, visible through cuticle; internal genitalia as in Figs 76View Figs 72 – 76, 91–92.

Natural history

All specimens were found under large dead leaves on the ground. They hung in domed sheet webs closely attached to the leaf surface. They barely moved when disturbed and were not seen to vibrate.


Known from type locality in Sumatra only ( Fig. 1View Fig. 1).


Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"


USA, Florida, Gainesville, University of Florida, Florida Museum of Natural History, Allyn Museum