Exocelina bacchusi herzogensis Shaverdo & Balke

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

publication ID

https://dx.doi.org/10.3897/zookeys.878.37403

publication LSID

lsid:zoobank.org:pub:192214DE-1D38-467B-A577-ECD16EC5EAB5

persistent identifier

https://treatment.plazi.org/id/BB967394-B81D-56C7-A135-25136A9E8C64

treatment provided by

ZooKeys by Pensoft

scientific name

Exocelina bacchusi herzogensis Shaverdo & Balke
status

ssp. nov.

6a. Exocelina bacchusi herzogensis Shaverdo & Balke ssp. nov. Figs 17 View Figures 12–17 , 23 View Figures 22, 23

Exocelina undescribed sp. MB1383: Toussaint et al. 2014: supplementary figs 1-4, tab. 2; Toussaint et al. 2015: supplementary figs S1, S2, tab. S3, and information S5, S6.

Type locality.

Papua New Guinea: Central Province, Woitape, 08°33.17'S, 147°15.48'E, 1500 m a.s.l.

Type material.

Holotype: male "Papua New Guinea: Central, Woitape, 1500m, i.2008, [08°] 33.178S 147.15.481E, Posman (PNG 167)", "DNA M.Balke 3401" [green] (ZSM). Paratypes: 1 male, 2 females with the same labels as the holotype (NHMW, ZSM). 1 male "Papua New Guinea: Morobe, Wagau, Herzog Mts., 1150m, 19.xi.2006, 06.51.067S 146.48.068E, Balke & Kinibel (PNG 102)", "DNA M.Balke 1383" [green] (ZSM).

Description.

Body size and form: Beetle small: TL-H 3.4-3.6 mm, TL 3.8-4.0 mm, MW 1.85-2.0 mm (holotype: TL-H 3.6 mm, TL 4.0 mm, MW 2.0 mm), with oblong-oval habitus.

Colouration: Yellow reddish to brown. Head reddish brown to brown, dark brown posterior to eyes. Pronotum yellowish reddish, with small dark area on disc or brown, with paler sides. Elytra yellow reddish to brown. Head appendages and legs proximally yellowish, legs distally darker, reddish brown ( Fig. 17 View Figures 12–17 ). Teneral specimen yellowish.

Surface sculpture: Shiny dorsally, with very fine punctation and weakly impressed microreticulation. Elytral punctation and microreticulation finer then in nominotypical subspecies. Elytral punctation usually invisible.

Structures: Pronotum with lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct bead and few setae laterally. Abdominal ventrite 6 slightly truncate.

Male: Protarsomere 4 with anterolateral seta rather long and thing, evenly curved, smaller than more laterally situated large seta. Protarsomere 5 ventrally with anterior band of more than 60 and posterior row of ten relatively long setae ( Fig. 23D View Figures 22, 23 ). Abdominal ventrite 6 with 6-8 lateral striae on each side. Median lobe simple, evenly tapering towards apex in lateral and ventral views; in lateral view, apex elongate, thin, with slightly enlarged, rounded tip ( Fig. 23A, B View Figures 22, 23 ). Paramere as in Fig. 23C View Figures 22, 23 .

Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.

Variability.

Colouration of the specimens from Woitape distinctly paler, yellowish; the specimen form Wagau much darker, brown.

Affinities.

From the nominotypical subspecies, it can be distinguished by shinier dorsal surface, shorter setae of male protarsomere 4, and by apex of the median lone elongate, thinner, with slightly enlarged tip. The further study is necessary to confirm the status of this taxon, which seems to replace the nominotypical subspecies in the Papuan Peninsula.

Distribution.

Papua New Guinea: Morobe and Central Provinces ( Fig. 25 View Figure 25 ).

Etymology.

The subspecies is named after Herzog Mts., where the subspecies was the first time discovered. The name is an adjective in the nominative singular.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Dytiscidae

Genus

Exocelina