Ellimenistes raucus Borovec & Nakládal, 2019

Borovec, Roman & Nakládal, Oto, 2019, Ellimenistes humeralis Marshall, 1947 and its allies (Coleoptera: Curculionidae: Entiminae: Embrithini), Zootaxa 4563 (1), pp. 175-184 : 182-183

Ellimenistes raucus Borovec & Nakládal , sp. nov.

( Figs 6, 9, 12, 15, 19 , 23, 29 ) http://zoobank.org/ urn:lsid:zoobank.org:act:93C2DAC3-BB1F-4FD3-855A-1CC89A870030

Type locality. South Africa, Eastern Cape, Silaka Forest Reserve .

Type material. Holotype: ♂, ‘ S. Afr. [ South Africa], Transkei [Eastern Cape], Silaka For. Reserve , 31.33 S– 29.30 E, 24.11.1987; E-Y: 2533, indig. forest litter, leg. Endrödy-Younga’ ( TMSA) . Paratypes: 1 ♂ 1 ♀, the same data as holoytpe ( TMSA) .

Description. Body length 3.72–4.16 mm, holotype 4.16 mm. As this newly described species is very similar to E. janaki sp. nov., with dorsal habitus as in Fig. 2 , the following description only provides characters differing in these species:

Rostrum ( Figs 6, 9 ) moderately longer, 1.23–1.28× wider than long. Scrobes in profile shorter than distance between scrobe and eye.

Antennal funicle with segment 1 the largest, conical, 1.5–1.6× longer than wide and 1.3–1.4× longer than segment 2, which is 1.5–1.6× longer than wide; segments 3–7 1.1× longer than wide.

Pronotum (as in Fig. 2 ) 1.39–1.47× wider than long.

Elytra (as in Fig. 2 ) long oval, 1.26–1.29× longer than wide, odd intervals indistinctly more elevated than even ones.

Penis ( Fig. 12 ) moderately long, slender, subparallel-sided, slightly widest at apical quarter; apex regularly tapered apicad with slightly concave sides and narrowly rounded tip; in profile with ventral side almost straight and dorsal side convex, apex lengthened, rounded at a tip. Tegmen ( Fig. 15 ) with slender ring and long, pointed parameres, almost as long as diameter of ring, connected at their basal half and forming here short basal plate.

Spermatheca ( Fig. 19 ) with short and wide, curved, pointed cornu; corpus small; ramus tube-shaped, more than twice longer than wide, at midlength distinctly curved downwards, not pressed to corpus; nodulus directed upwards, almost straight, tube shaped, 3.5× longer than wide at midlength, widest at base and slightly evenly tapered apicad, slightly longer than cornu. Sternite VIII ( Fig. 23 ) with plate subtriangular, 1.8× longer than wide, with straight sides. Gonocoxites ( Fig. 29 ) flat, evenly tapered apicad with long slender apical styli with setae.

Biology. The type material was collected from native forest litter.

Etymology. The Latin name, meaning “rough, coarse”, refers to not smooth aspect of body vestiture.

Differential diagnosis. Ellimenistes raucus sp. nov. is very similar to E. janaki sp. nov., both species can be distinguished by the characters given below in the key.

Figures

FIGURES 4–19. Structural details of Ellimenistes Boheman. Figs 4–6, head with rostrum, dorsal view: 4. E. humeralis Marshall; 5. E. janaki sp. n.; 6. E. raucus sp. n. Figs 7–9, head with rostrum, lateral view: 7. E. humeralis Marshall; 8. E. janaki sp. n.; 9. E. raucus sp. n. Figs 10–12, penis, ventral and lateral view: 10. E. janaki sp. n.; 11. E. marshalli sp. n.; 12. E. raucus sp. n. Figs 13–15, tegmen, dorsal view: 13. E. janaki sp. n.; 14. E. marshalli sp. n.; 15. E. raucus sp. n. Figs 16–19, spermatheca: 16. E. humeralis Marshall; 17. E. janaki sp. n.; 18. E. marshalli sp. n.; 19. E. raucus sp. n.

FIGURES 20–29. Structural details of Ellimenistes Boheman, Cycliscus Schoenherr and Cosmorhinus Schoenherr. Figs 20– 25, sternite VIII of female, dorsal view: 20. Ellimenistes humeralis Marshall; 21. E. janaki sp. n.; 22. E. marshalli sp. n.; 23. E. raucus sp. n.; 24. Cycliscus caecicollis Boheman; 25. Cosmorhinus cervinus Boheman. Figs 26–29, ovipositor, dorsal view: 26. Ellimenistes humeralis Marshall; 27. E. janaki sp. n.; 28. E. marshalli sp. n.; 29. E. raucus sp. n.

FIGURES 1–3. Dorsal habitus of Ellimenistes Boheman. 1. E. humeralis Marshall; 2. E. janaki sp. n.; 3. E. marshalli sp. n.