Sparganothoides carycrosana Kruse and Powell, 2009

Kruse, James J. & Powell, Jerry A., 2009, Systematics of Sparganothoides Lambert and Powell, 1986 (Lepidoptera: Tortricidae: Sparganothini), Zootaxa 2150 (1), pp. 1-78 : 30-31

10. Sparganothoides carycrosana Kruse and Powell , new species

Figs. 16 , 70

Diagnosis. This species is superficially most similar to S. teratana but lacks the patch of greenish yellow scales characteristic of the latter. The uncus has a strongly bilobed apex in S. carycrosana , whereas it is more rounded apically in S. teratana . The genitalia of S. carycrosana are very similar to those of S. plemmelana , but the two species are dissimilar in coloration.

Description. Male. Head: Frons pale brown to brownish gray, produced; vertex roughened, pale brown to brownish gray, occasionally with a gray median stripe. Labial palpus short, about twice as long as wide, light brown mesally, brown to brownish gray or brownish orange laterally. Antennal scaling brown. Thorax: Dorsum smooth scaled, light brown to grayish brown. Forewing length 8.3–10.3 mm (= 9.4; n = 10). Forewing costal fold short, less than one-half wing length; forewing ground color brown to brownish orange with dark brown to black rows of transverse strigulae throughout; light brown basal fascia with some orange suffusion in some specimens; brownish orange to dark brown median fascia distinct basally, extending from basal fascia, wide laterally and becoming diffuse, indistinctly linking to orange brown to dark brown subterminal fascia. Fringe yellowish white to brownish orange. Hindwing with irregular gray scaling, some specimens with dark gray transverse striae throughout. Abdomen: Genitalia ( Fig. 16 ; slide #5546; EME; Mexico, Sinaloa, 2 mi SW Potrerillos; n = 14) with uncus long, curved, apex dorso-ventrally flattened, broad, strongly bilobed, U-shaped, with long setae dorsally, broad patch of spinules ventrally; tegumen raised into an exaggerated oval process at base of uncus, laterally lobed near base of socius/gnathos arms; socius subtriangular to ovately rounded posteriorly, secondary arms long, slender, abruptly curved before enlarged apices, enlarged apices asymmetrically widened, splayed in fanlike configuration; transtilla sclerotized, straight, spines short, numerous over most of posterior margin, anterior lobed process reinforced with invagination at middle; valva subpentangular without sclerotized crease, costa straight in basal half, abruptly angled toward apex; sacculus slightly concave; pulvinus absent; phallus pistol-shaped, aedeagus tapered, attenuate apically, shorter than phallobase, attached to juxta by a strongly sclerotized process, phallobase elongate with a small rounded bulb; cornuti in a dense clump of> 30, with a minute spine near base.

Female. Unknown.

Type material. Holotype: Male : MEXICO: SINALOA: 2 mi SW Potrerillos, 4200', 12.viii.1986, J. Brown & J. Powell ( EME).

Paratypes (22♂). COSTA RICA: GUANACASTE: Rincon National Park, Mirador , Ad. , 900 m, 29.iii.1984 (1♂), D. Janzen ( INBio) . Estación Cacao, lado suroeste Volcán Cacao , 1100 m, 8–30.vi.1991 (1♂), C. Chaves ( INBio) , 2–9.v.1992 (1♂), C. Moraga & P. Rios ( INBio) . PUNTARENAS: Estación Biol., Las Alturas , 12 km NE San Vito, 1550 m, 22–24.i.1993 (2♂), J. Powell ( EME) . GUATEMALA: Chejel , [no date] (3♂), W. Schaus & Barnes ( NMNH) . Cayuga , [no date] (1♂), W. Schaus ( NMNH) . Baleu, Mpio. San Cristóbal, Verapaz , Alta Verapaz, 1350 m, 15.viii.1985 (3♂), E. Welling ( EME) . MEXICO: SINALOA: 2 mi SW Potrerillos , 4200', 12.viii.1986 (1♂), J. Brown & J. Powell ( EME) . TAMAULIPAS: El Encino, 250 m, 4–13.viii.1988 (1♂), V. Becker ( VBC) . Gómez Farias , 1200 m, 26.v.1997 (1♂), V. Becker ( VBC) . VERACRUZ: Fortín de las Flores , 24.vii.1966 (1♂), O. Flint & Ortiz ( NMNH) . Fortín de las Flores , 1010 m, 7–12.vii.1974 (4♂), J. Chemsak, E. Linsley & J. Powell ( EME, NMNH) . Río Metlec Canyon, NW of Fortín de las Flores , 6.vii.1974 (1♂), J. Chemsak & J. Powell ( EME) . Cordoba , 930 m, 11.vii.1974 (1♂), J. Chemsak & J. Powell ( EME) .

Immature stages. Unknown.

Biology. Adults have been captured from January through June in Costa Rica and from May through August in Guatemala and Mexico, probably reflecting multiple broods.

Distribution. The species ranges from Sinaloa and Tamaulipas to Veracruz, Mexico, south through Guatemala to Costa Rica.

Etymology. The name is derived from the Greek “karykrous” (= nut-brown).

Figures

FIGURES 13–18. Male genitalia of Sparganothoides; valvae spread, aedeagus removed. 13. S. audentiana. 14. S. canitites. 15. S. teratana. 16. S. carycrosana. 17. S. plemmelana. 18. S. lugens.

FIGURES 61–78. Adults of Sparganothoides. 61. S. hydeana (paratype: USA: NM: Santa Fe County, Little Tesuque Canon, 9200', 27.vii – 10.viii.1932, A. Klots). 62. S. machimiana (USA: AZ: Santa Cruz County, 7 mi. SE Nogales, 1.viii.1991, P. Opler, Y. Hsu and D. Rubinoff). 63 S. laderana (holotype). 64. S. amitana (paratype). 65. S. calthograptana (holotype). 66. S. coloratana (holotype). 67. S. audentiana (holotype). 68. S. canities (paratype). 69. S. teratana (Panama: Cerro Campana, 11–14.vii.1967, O. Flint, Jr.). 70. S. carycrosana (holotype). 71. S. plemmelana (holotype). 72. S. lugens (Mexico: Veracruz: 1 mi. W Fortin, 25.xii.1963 as larva on miscellaneous leaves, emerged 20.ii.1964, C. O'Brien). 73. S. castanea (holotype). 74. S. vinolenta (Mexico: Distrito Federal: La Venta, 22.v.1938, T. Escalante). 75. S. ocrisana (paratype, male: Costa Rica: Puntarenas, Monteverde, 1350–1400m, 22–24.vii.1990, J. Powell, JAP lot 90G14, emerged 6.xi.1990). 76. S. ocrisana (paratype, female: Costa Rica: Puntarenas, Monteverde, 1350–1400m, 22–24.vii.1990, J. Powell). 77. S. xenopsana (holotype). 78. S. umbosana (holotype).