Croton minimimarginiglandulosus Radcl.-Sm., Gen. Croton Madag. Comoro 18. 2016.

Berry, Paul E., Kainulainen, Kent & Ee, Benjamin W. van, 2017, A Nomenclator of Croton (Euphorbiaceae) in Madagascar, the Comoros Archipelago, and the Mascarene Islands, PhytoKeys 90, pp. 1-87 : 48-49

93. Croton minimimarginiglandulosus Radcl.-Sm., Gen. Croton Madag. Comoro 18. 2016.

Croton fianarantsoae var. ambremontanus Radcl.-Sm., Gen. Croton Madag. Comoro 153. 2016, syn. nov. Type. Madagascar. Prov. Antsiranana: National Park of Montagne d’Ambre, path from station to Bianamalao, 12°32'S, 49°10'E, 1120 m, 4 Jun 1989, B. Du Puy, D. Du Puy, & G. Guy 218 (holotype: K!; isotype: P [P00060699]!).

Croton ivohibensis var. aesculops Radcl.-Sm., Gen. Croton Madag. Comoro 148. 2016, syn. nov. Type. Madagascar. Prov. Antsiranana: Reserve Spéciale d’Ankarana, 12°54'43"S, 49°06'39"E, 19 Feb 1994, M. Andrianarisata et al. 43 (holotype: K!; isotypes: MO!, P [P00433266]!).

Croton ivohibensis var. ankaranaensis Radcl.-Sm., Gen. Croton Madag. Comoro 148. 2016, syn. nov. Type. Madagascar. Prov. Antsiranana: Reserve Naturelle Intégrale d’Ankarana, 12°51'S, 49°05'E, 18 May 1987, M. Nicoll & J.P. Abraham 684 (holotype: K!; isotypes: MICH [MICH1514788]!, MO!, P [P00133271]!).

Croton thouarsianus var. macrocalyx Radcl.-Sm., Gen. Croton Madag. Comoro 183. 2016, syn. nov. Type. Madagascar. Antsiranana: Ankarana Reserve, Tsingy area, 12°54'42"S, 49°06'42"E, 22 May 1993, C.C.H. Jongkind & S. Rapanarivo 960 (holotype: K!; isotypes: MICH [MICH1514787]!, MO!, P [P00433151]!).

Type.

Madagascar. Prov. Antsiranana: au sud d’Antsiranana, près de Joffreville dans le Parc National de Montagne d'Ambre , 12°27'S, 49°13'E, 3-10 Aug 1993, O. Andrianantoanina & Rocsceohclher 283 (holotype: K!; isotype: MO!) .

Habit and distribution.

Shrubs; northern Madagascar (Antsiranana).

Notes.

This species is restricted to Montagne d’Ambre and tsingy habitats in the Ankarana Special Reserve. It has somewhat anisophyllous terminal leaves like C. thouarsianus , but the calyx of the pistillate flowers is much larger and more accrescent in fruit, with a long pedicel. Plants vary considerably in pubescence, with some hirsute stems in the type of C. thouarsianus var. macrocalyx . Quoting from Radcliffe-Smith (2016), "The admittedly rather cumbersome name, of 10 syllables & 24 letters, can be justified on the grounds of the distinctive minute ogivaliform marginal glands to be found on the otherwise entire leaf-margins of this species. The name is not occupied in the genus." No surprise there!