Craspedophorus tetrastigma milzi Duvivier, 1891,

Häckel, Martin, 2017, A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 3. Revision of the Craspedophorus strachani and C. brevicollis groups (Coleoptera: Carabidae), Zootaxa 4330 (1), pp. 1-67: 29

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Craspedophorus tetrastigma milzi Duvivier, 1891

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17b. Craspedophorus tetrastigma milzi Duvivier, 1891  new status

( Plate 3View PLATE 3, Figs 24–27; Plate 4View PLATE 4, Fig. 30, Plate 12View PLATE 12, Fig. 99)

Craspdophorus milzi Duvivier, 1891  : ccclxxvi (type locality “Djabir-Bandja [=Bondo (Bakongo), Orientale (Bas-Uele) Province, Democratic Republic of the Congo]”). Basilewsky 1987: 195, Lorenz 2005: 321. Häckel and Farkač 2012: 82 [with erroneous locality].

Craspedophorus strachani milzi Basilewsky 1951b: 206  .

Eudema  (ou Craspedophorus  ) strachani  var. nov  . depressus Alluaud, 1929: 89 (type locality “Bas-Uelé [=Orientale Province, Democratic Republic of the Congo]”). Craspedophorus strachani depressus  (? milzi  ) Burgeon 1930b: 158. Craspedophorus strachani milzi  (depressus) Burgeon 1935b: 180. Craspedophorus strachani depressus Lorenz 2005: 321  . Häckel and Farkač 2012: 83. syn. nov.

Type material. 1♀ labeled as holotype Craspedophorus strachani depressus Alluaud, 1929  : “ Strachani / v. depressus  / Type Alluaud [handwritten in black]/ Alluaud, det. 192 [printed in red on white red circumscribed label]// R. dét. [printed in black]/ W [handwritten in black]/ 1454 [printed in black on white label]// depressus / Alld. / (= Milz i / Duv.) [handwritten in black on white label]// Craspedophorus  / strachani-depressus [handwritten in black] dét. Ch. Alluaud 1927 [printed in black on white label]// [DataMatrix] RMCA Ent / 0 0 0 0 20052 [printed on white label]” ( Plate 3View PLATE 3, Fig. 25, MRAC). 1♂ labeled as paratype C. s. depressus Alluaud, 1929  : “région du Bas- Uelé / 7–18 nov. 1925 / Prince Léopold [printed in black on white label]// Don du / Musée royal / de Belgique [handwritten in black on white label]// ex Typis [printed in red on white label]// depressus [handwritten in black on white label]// Muséum Paris / Coll. Ch. Alluaud [printed in black on blue label]// Craspedophorus  / strachanidepressus [handwritten in black] dét. Ch. Alluaud 1927 [printed in black on white label] ( Plate 3View PLATE 3, Fig. 26, Plate 4View PLATE 4, Fig. 30, MNHN), 1♂ same labeled except: “Paratype [printed in black on red label] (MRAC).

Additional material examined. Central African Republic. 1 Ƌ: “Yalinga /( Oubangui ) [Haute Kotto Prefecture]” / G.Le Testu // cf. Milzi Duv.  / t. Burgeon // Muséum Paris / Coll. Ch. Alluaud ” ( Plate 3View PLATE 3, Fig. 24, MNHN)  . DR Congo: Équateur. 1♂: “Région Lisala”; 1♀: “Ubangi: Molegbwe”; 1♀: “ M'Paka (terr. Libenge)” ( MRAC)  . Orientale. 1♂: “Ibembo, Congo Belge ” ( NMPC); 2♂, 2♀: “Bas-Uelé: Buta” ( Plate 3View PLATE 3, Fig. 27, Plate 12View PLATE 12, Fig. 99, MRAC, cMH)”; 1♂: “Uele: Lakulu”; 2♀: “Uele: Monga” ( MRAC). 

Note. This species is based on specimens labeled “Djabir-Bandja”. Bondo (Djabir) is a place on Uele River with a very small population in the region of Orientale, DR Congo. I have not seen the type deposited in Brussels, my opinions are based on the profound examination by Burgeon (1930b: 158) and Basilewsky (1951b: 206). Description (in part, see Duvivier 1891: ccclxxvi). “Length 19 mm, width 8 mm. Black, glossy, almost parallelsided; pronotum quite similar to that in C. tetrastigma ( Chaudoir, 1850)  , but rugate, punctured, lateral margins posteriorly more elevated (wider at midlength), more narrowing anteriorly; elytra almost parallel-sided, moderately flattened, black, glossy, punctured, striated, intervals rugate, punctured, each with two red maculae; humeral macula near base, with margin, obliqued anteriorly, reaching from III to VII interval, limited to anterior third, preapical macula near apex, with anterior margin in its apical third, hind margin obliqued posteriorly, reaching from IV to VIII interval, with small macular spot on III interval, maculae not regularly marginated, except anterior margin in humeral macula, which is almost flat. Body and legs black” [from French]. Later C. strachani depressus Alluaud, 1929  , was based on three specimens collected in Bas-Uele District, Orientale Province, DR Congo. Description (in part, see Alluaud 1929: 89). “...represent one well distinct subspecies (I do not establish one new different species, I prefer to treat it as a variant of C. strachani  ). Body less convex posteriorly, markedly depressed on disc; yellow humeral maculae with more equalised spots, maculae often more serrate, because of mostly denticulated posterior margins; preapical maculae, without more equalised spots, but as well more serrate. Elytral margins moderately more rounded, creating more ovoid back of the body, less distinctly anteriorly as well as posteriorly” [from French]. Basilewsky added (1951b: 206). “ C. Strachani  Hope is a rather variable species, yet I do not hesitate to separate this series of specimens which, as stated above, are quite similar to one another because of a certain variation in colourization which distinguishes west African from central African specimens at first sight. On the other hand, constant differences between true Strachani from West Africa and Milzi Duvivier  (= depressus Alluaud) race from  the North of Congo are not significant, yet present in all Congoan specimens. Therefore I consider all specimens from the French Congo and from the North of The Belgian Congo one subspecies ssp. Milzi, not greatly different from true Strachani ” [from French].

Distribution. Central African Republic, Democratic Congo: Équateur, Orientale Provinces.


Museum National d'Histoire Naturelle


Mus�e Royal de l�Afrique Centrale


National Museum Prague














Craspedophorus tetrastigma milzi Duvivier, 1891

Häckel, Martin 2017

Craspdophorus milzi

Hackel 2012: 82
Lorenz 2005: 321
Basilewsky 1987: 195


Hackel 2012: 83
Lorenz 2005: 321
Burgeon 1935: 180
Burgeon 1930: 158
Alluaud 1929: 89

Craspedophorus strachani milzi

Basilewsky 1951: 206