Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomics-based taxonomic rearrangement of Achlyodini and Carcharodini (Lepidoptera: Hesperiidae: Pyrginae), Insecta Mundi 2023 (16), pp. 1-33 : 8

publication ID

https://doi.org/ 10.5281/zenodo.10621955

persistent identifier

https://treatment.plazi.org/id/BF04E441-FFD5-2A04-B996-43FD58DFFC74

treatment provided by

Felipe

scientific name

Grishin
status

subgen. nov.

Tuberna Grishin , new subgenus

http://zoobank.org/ 7C63B146-7D14-42AC-B8EC-071C103D39D4

Type species. Pythonides contubernalis Mabille, 1883 View in CoL .

Definition. The newly expanded genus Quadrus Lindsey, 1925 View in CoL (type species Papilio cerialis Stoll, 1782 ) can be divided into eight clades at the subgenus level in the tree ( Fig. 1 View Figure 1 ). In addition to the nominotypical subgenus, two of these clades correspond to traditional subgenera, previously treated as genera: Ouleus Lindsey, 1925 View in CoL (type species Achlyodes fridericus Geyer, 1832 View in CoL ) and Zera Evans, 1953 View in CoL (type species Achlyodes zera Butler, 1870 View in CoL ), and five clades represent new subgenera. One of these new subgenera is sister to all others except Trifa new subgenus and keys to or E.39.4a or E. 41.1 in Evans (1953) and is distinguished from its relatives by the following characters: dorsal hindwing either with 2 bright shiny blue bands, or primarily blue with dark veins and central white streak; uncus undivided, stem-shaped, rounded at its end, with parallel sides in dorsal view, ampulla with projection (or a hump), harpe longer than valva, either with concave dorsal margin or rounded, nearly oval, merged with valva and ampulla with a small hump. In DNA, a combination of the following nuclear genomic base pairs is diagnostic: aly 2116.6.1:T200C, aly 2116.6.1:C326T, aly1475.25.1:A1840C, aly1475.25.1:T1200A, aly1838.49.3:G435T.

Etymology. The name is a feminine noun in the nominative singular, formed from the type species name: [con] Tuberna [lis].

Species included. Pythonides contubernalis Mabille, 1883 , Pythonides deyrollei Mabille, 1877 , and Leucochitonea lancea Hewitson, 1868 .

Parent taxon. Genus Quadrus Lindsey, 1925 .

Darwin Core Archive (for parent article) View in SIBiLS Plain XML RDF