DALMANNIINAE HENDEL, 1916

Gibson, Joel F. & Skevington, Jeffrey H., 2013, Phylogeny and taxonomic revision of all genera of Conopidae (Diptera) based on morphological data, Zoological Journal of the Linnean Society 167 (1), pp. 43-81 : 66-67

DALMANNIINAE HENDEL, 1916

Type genus Dalmannia Robineau-Desvoidy, 1830: 248 . Included genera: Baruerizodion Papavero, 1970: 121 , Dalmannia . Uncontroverted morphological autapomorphies: female abdominal sternite and tergite 6 separate (88); female abdominal segment 7 compressed laterally along entire length (97). Ocellar bristles are absent in all members of Dalmanniinae (14); this character state is also observed in Pyrgotidae , Syrphidae , Conopinae , and some species of Stylogaster . Postocellar bristles are absent in all members of Dalmanniinae (15); this character state is also observed in Pyrgotidae , Syrphidae , and Conopinae .

Gibson et al. (2012) recover Dalmanniinae as monophyletic with the same set of apomorphic character states, but only include representatives of Dalmannia . They include the following additional apomorphic character states: phallus coiled and setose; and posterior surstyli absent. Hendel’s (1916) original mention of Dalmanniinae does not specify included genera and chracter states. It only states that Dalmanniinae possesses an unbroken costa and ‘[other primitive characteristics]’.

Zimina (1960) describes the tribe Dalmanniini with the following key characteristics: female with long, curved ‘ovipositor’; phallus elongate and ‘bandshaped’; veins Sc and R 1 ending not near one another; cell cu p short; lower pleural sclerites bare. According to the present analysis, the male and female terminalia characters are both autapomorphies of Dalmanniinae . For the wing and pleural characters, Dalmanniinae retains the condition plesiomorphic to Conopidae . The presence of an ‘ovipositor’ has previously been used to define Dalmanniinae , and in the present analysis, modification to female abdominal segment 7 is an autapomorphy of the subfamily. Superficially similar, yet distinct, modifications to the female terminalia have occurred in other subfamilies (e.g. Paramyopa , Parazodion , and Tanyconops ).

While all other authors have afforded Dalmanniinae subfamily status, it was described by Zimina (1960) as a tribe of Myopinae . According to the present analysis, in addition to its apomorphic character states, Dalmanniinae can be distinguished from Myopinae by a broad basisternum, unfused veins Sc and R 1, absence of spicules on female abdominal sternites 5 and 6, absence of spicules on male abdominal sternite 5, narrow male syntergosternite 7 + 8, and absence of male abdominal sternite 8.

Species examined – Baruerizodion steyskali ; Dalmannia aculeata ; Dalmannia nigriceps ; Dalmannia vitiosa .