Eupines (Byraxis) anisarthra (Broun, 1914)

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 11-12

Eupines (Byraxis) anisarthra (Broun, 1914)

Fig. 4 ; map Fig. 49

Byraxis anisarthra Broun, 1914b: 169 . Hudson 1923: 364, 1934: 185.

Eupines (Byraxis) anisarthra : combination and subgeneric assignment by Nomura and Leschen 2006: 254; Maddison 2010: 417; Shen and Leschen 2019: 259 .

Type locality: [ MC] Broken River .

Type depository: BMNH LT ♂, 1 PLT .

Diagnosis: Head with vertexal foveae punctiform; antennae 10-segmented with 2-segmented club; A4, A6, A7, A8 and A10 unmodified, A4 slightly asymmetrical, A5 large and asymmetrical, wider than A4 at middle and longer than A3 + A4, outer margin flat, A9 large, about 4 × longer than A8, A10 conical, about 1.6 × longer than A9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, about one-fourth of ventral length. Protrochanters with barely visible ventral spine, shorter than one-tenth of protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.51–1.63 mm; body colour reddish brown to dark brown with lighter coloured elytra and legs. Head about as long as wide; frons narrowed at end, slightly shorter than half width of head at eye level; eyes well developed, with about 34 facets; vertexal foveae punctiform; lacking antennal tubercles and postantennal notches; postantennal cavities small, slightly shorter than half length of antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented club; A4, A6, A7, A8 and A10 unmodified, A4 slightly asymmetrical, A5 large and asymmetrical, wider than A4 at middle and longer than A3 + A4, outer margin flat, A9 large, about 4 × longer than A8, A10 conical, about 1.6 × longer than A9. Pronotum wider than long, slightly wider than head at eye level; Elytra slightly wider than long, widest near middle; humeral calli slightly prominent; sutural stria distinctive at base and vaguely visible near apex. Abdomen with tergite 1 about 1.5 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved apical setae; ventrite 6 with pair of median struts, about one-fourth of ventral length. Legs with protrochanters modified with barely visible ventral spine, shorter than one-tenth of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.28 mm long; with elongate and thin parameres, curved downwards and twisted at apex, with two long subapical setae; median lobe with two long sclerites, slightly wider than parameres in dorsal view, left one curved inward subapically, right one bent abruptly.

Comments: Eupines (B.) anisarthra is similar to E. (B.) minuta by having A5 and A9 similarly modified and can be separated by the unmodified mesotibiae (a small apical spine is present in E. (B.) minuta ; Fig. 31E ).

Distribution: South Island. MC, CO.

Additional material examined: 3 ♂♂: CO: 2, Carrick Range, Watts Rock, 1380 m, 31/Oct/1979, J.C. Watt, moss, 79/133; 1, JNPC, Old Man Range, Headwaters, Obelisk Ck., 1380 m, 13/Jan/07. Associated females (2 ♀♀): 2, CO: Carrick Range, Watts Rock, 1380 m, 31/Oct/1979, J.C. Watt, moss, 79/133.

Figures

FIGURE 4. Diagnostic characters of E. (B.) anisarthra (Broun, 1914). A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 31. Diagnostic characters of E. (B.) mayae sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 49. Geographic distributions of E. (B.) anisarthra (Broun, 1914), E. (B.) bisulcifrons (Broun, 1914), E. (B.) brevis sp. n., E. (B.) caesta sp. n., E. (B.) carinata sp. n., E. (B.) coalita sp. n., E. (B.) complector sp. n. and E. (B.) conspicua (Broun, 1893) in New Zealand.