Actinopus pampa, Ríos-Tamayo & Goloboff, 2018

Ríos-Tamayo, D. & Goloboff, P. A., 2018, Taxonomic Revision And Morphology Of The Trapdoor Spider Genus Actinopus (Mygalomorphae: Actinopodidae) In Argentina, Bulletin of the American Museum of Natural History 2018 (419), pp. 1-84 : 55-59

Actinopus pampa , sp. nov.

Figures 27A–J, 28A–E, 45

TYPE MATERIAL: HOLOTYPE ♂ (MACN-Ar.

28419): La Pampa: Santa Rosa [36°37′ S 64°17′

W], Apr.-May. 1967, Aravena, 1♂. PARATYPE: 1♀ (MACN-Ar 36623), Parque Nacional “Lihuél Calel” [37°59′ S 65°35′ W], Jan. 1983, P. Goloboff.

ETYMOLOGY: The specific name is a noun in apposition that refers to the type locality of the species.

DIAGNOSIS: Females of A. pampa can be recognized by their simple and very small spermathecae (fig. 28E), wide, subquadrate, with a very small apical lobe, similar to that of A. ramirezi from which it differs by its more rounded fovea (fig. 28A). Males can be distinguished by having a copulatory bulb with a rounded shape and stylized embolus (similar to that of A. argenteus and A. longipalpis ) with a ventral depression in the tegulum (fig. 27B). Differs from all other species (except from A. argenteus ) in the silvery color on the abdomen dorsum; differs from A. argenteus in having fewer retrolateral spines on tibia II.

DESCRIPTION: Holotype male (MACN-Ar. 28419, fig. 27A–J): Total length: 17.76. Carapace (fig. 27A): length 6.60, width 6.24. Cephalic region: Length 3.48, 4.68 width; clypeus short with one large bristle and 15 small ones on both sides; 6 bristles between AME-ALE, 9 small ones between PME and 3–4 behind each PME-PLE. 2–3 small bristles on each side of the OQ, the central larger. Fovea: length 0.96, width 2.64. Eyes, diameters and interdistances: AME 0.40 (large), ALE 0.48, PME 0.28, PLE 0.40, OQ length 1.28, anterior width 3.40, posterior width 3.20; AME-ALE 0.76, PME-PLE 0.16, AME- PME 0.48, ALE-PLE 0.60, AME-AME 0.20. Chelicerae: Length 3.24, width 1.86; cheliceral apex with 15 recumbent and elongated bristles that cover the base of the fang. Cheliceral prolongation without cusps; furrow with 6 large promarginal teeth and 6 retromarginal (a small denticle between 5th-6th) and 15 denticles, near the promarginal teeth. Labium: 1.26 length, 1.20 width. Maxillae: width 1.92, anterior length 2.10, posterior length 2.40. Sternum (fig. 27J): Center flat; length 4.62, maximum width 3.60, minimum width 1.38. Postlabial sigillum shallow, posterior sigilla well defined and shallow, separated from margin. Abdomen (fig. 27C, F): length 7.20, covered by small bristles, thicker on dorsum. PMS: length 0.56; PLS: length of basal:medial:apical segments 0.72:0.28:0.20. Spigots: 1 large, 25 small on PMS; PLS, numbers of large(small) spigots on basal:medial:apical article 2(30): (the other articles deformed, could not be observed). Lengths of legs and palp: I: 5.76, 2.70, 3.24, 4.14, 2.10, 17.94. II: 5.70, 2.70, 3.24, 4.32, 2.10, 18.06. III: 4.80, 2.88, 2.70, 4.80, 2.52, 17.70. IV: 6.00, 3.00,

4.86, 5.40, 2.76, 22.02. Palp: 5.34, 2.64, 4.50, –, 1.50, 13.98.

Chaetotaxy: All femora: 0. Patellae: I-II, 0; III, 40 P SUP-D ANT +10 on the margin, 1-2-1-2- 1-1 D, 7 R on the margin; IV, 50 P SUP-D ANT. Tibiae: I, 0; II, 1-1-1-2 R INF, 1-1 V POST; III, 1 P, with an apical crown with 18 thorns; IV, 2-1 P INF (1:2 A). Metatarsi: I, 2-1-2-3 V; II, 1-1-1-1 R INF, 1-2 V ANT (1:4 A), 1-1-1 v post; III, 1-1- 2-1-2 P INF, 1 R A, 1 V A POST; IV, 2-1-2-2-3-2 V. Tarsi: I, 1-1-1-1-1 P INF, 2-2-1-3 R INF; II, 1-1-1-1-1-1 P INF, 1-5-3-5 R INF; III, 1-2-2-2- 1-4 P INF, 1-1 R INF; IV, 1-3-3-4-3-4 P INF, 1-1-2-1-2-1 R INF. Total number of retrolateral spines on tibiae I and II (fig. 27E): 0 and 5 respectively. Palp: Tibia short and thickened (width/length: 0.36±0.01) (fig. 27D). Bulb (fig. 27G–I) with a rounded tegulum, BTA and ATA poorly developed. Pseudoscopula: Tarsi: I-II sparse (uniformly distributed); III-IV, dense and uniform. Metatarsi: IV sparse and apical. Trichobothria: Tibiae: I-III 6-10; IV 10-12. Metatarsi: I 17; II 17; III 14; IV 10. Tarsi: I-II 10; III 12; IV 8. Color: Carapace, cephalic region and

chelicerae dark, almost black reddish brown. Legs like carapace with lighter tibiae-metatarsitarsi, palp with lighter tibia. Sternum reddish, labium and maxillae darker. Abdomen dark brown, dorsum silver. Booklungs with dark markings. Spinnerets yellowish.

Paratype female (MACN-Ar 36623, fig. 28A– E): Total length: 15.60. Carapace (fig. 28A): length 5.10, width 4.62, with 16 bristles on each posterior margin. Cephalic region: length 3.06, width 3.90; clypeus with 35 bristles, central larger, chilum short with 16 bristles; 28 bristles between PME, 5–7 behind each PME-PLE dyad; 10–12 bristles on each side of the OQ, central larger. Fovea: length 0.90, width 2.10. Eyes, diameters and interdistances: AME 0.14, ALE 0.32, PME 0.16, PLE 0.28, AME-ALE 0.80, PME-PLE 0.08, AME-PME 0.48, ALE-PLE 0.52, AME-AME 0.24, ocular region length 0.96, anterior width 2.68, posterior width 2.40. Chelicerae: Length 2.34, width 2.34; chelicerae apex with 11 recumbent and elongated bristles covering the base of the fang. Rastellum with 11 marginal, blunt cusps, and 6 small dorsal ones. Fang groove with 6 promarginal teeth and 5 retromarginal ones and 12 denticles in the furrow, near the promargin. Labium: length 1.02, width 1.02, with 14 anterior cuspules. Maxillae: width 1.32, anterior length 1.56, posterior length 2.04, with +80 cuspules 1:3 V ANT, extending to the anterior face. Sternum (fig. 28D): Center flat; length 3.48, maximum width 3.00, minimum width 1.02. Postlabial sigillum shallow, posterior sigilla shallow and well defined, separated from margin. Abdomen: Globose, covered by dark fine bristles. PMS: length 0.40; PLS: length of basal:medial:apical segments 0.60:0.28:0.20. Spigots: 2 large, 25 small on PMS; PLS, numbers of large(small) spigots on basal:medial:apical article 4(30):8(40):2(50). Lengths of legs and palp: I: 2.94, 1.98, 1.26, 1.62, 0.78, 8.58. II: 3.00, 1.98, 1.32, 1.74, 0.78, 8.82. III: 3.00, 2.10, 0.66, 1.98, 0.78, 8.52. IV: 3.78, 2.16, 2.16, 2.88, 0.78, 11.76. Palp: 2.70, 1.80, 1.80, –, 1.92, 8.22.

Chaetotaxy: Femora: 0. Patellae: I-II, 0; III; 10 P SUP-D ANT (1:2 A) +12 on the margin, 19 R (1:2

A) +10 on the margin; IV, 56 P SUP-D ANT +5 near the edge; palp, 1-1-1 P, 1 V POST. Tibiae: I, 1-1-1 P A, 15 R INF, 1-1 V POST; II, 78 R INF, 1-1 v post; III, 3 P A, 1 D B, with an apical crown of 22 thorns and 3 apical (at the apex, anterior to the crown) 42 R; IV, 0; palp, 9 P INF, 20 R INF. Metatarsi: I, 16 P INF, 16 R INF, 1-1-2 V POST; II, 12 P INF, 10 R INF, 1-1-2 V POST; III, 2-1-2-3-2-1-1-2 D ANT, 33 on row D POST-R (more abundant R A), 3 V A; IV, 9 P (1:2 A), 1 D A ANT, 1 V A ANT. Tarsi: I, 9 P INF, 14 R INF, 2 V A; II, 13 P INF, 15 R INF, 2 V A; III, 23 R, 55 V-P; IV, 35 V-P. Total number of retrolateral spines on tibiae I and II (fig. 28B): 15 and 78 respectively. Trichobothria: Tibiae: I-II 4-7; III 5-9; IV 7-5. Metatarsi: I 12; II 14; III 11; IV 7. Tarsi: I 8; II 9; III 8; IV 4. Color: Carapace light reddish brown; clypeus with a light spot. Sternum reddish brown. Spinnerets yellowish. Abdomen light brown, booklungs with light markings. Legs like carapace.

DISTRIBUTION: Province of La Pampa.

ADDITIONAL MATERIAL EXAMINED: None.