Nocarodes

Ünal, Mustafa, 2016, Pamphagidae (Orthoptera: Acridoidea) from the Palaearctic Region: taxonomy, classification, keys to genera and a review of the tribe Nocarodeini I. Bolívar, Zootaxa 4206 (1), pp. 1-223 : 189-191

publication ID

https://doi.org/ 10.11646/zootaxa.4206.1.1

publication LSID

lsid:zoobank.org:pub:E1566C02-9987-4116-83AA-91D3D1DCF2FF

DOI

https://doi.org/10.5281/zenodo.5781160

persistent identifier

https://treatment.plazi.org/id/C24587A5-FF5A-4FEF-FF50-FA0042E5FAB7

treatment provided by

Plazi

scientific name

Nocarodes
status

 

Key to species groups, species and subspecies of Nocarodes View in CoL

1 Apical valves of penis regularly narrowing, pointed at apex; posterior margin in lateral view at most slightly concave; preapical part compressed antero-posteriorly ( Figs. 82 View FIGURES 69 – 82 a–89a).......................................................2

- Apical valves of penis distinctly swollen and widened in preapical part, blunt at apex; posterior margin in lateral view dis-

tinctly incised ( Figs. 90 View FIGURES 83 – 92 a, 775, 776)................................................. ( N. iranicus View in CoL sp. group)...16 2 Apical valves of penis small and very slender, their ventral margins turn to lateral sides, preapical part strongly compressed antero-posteriorly ( Figs. 82 View FIGURES 69 – 82 a–88a); body of variable size, but relatively slender ( Figs. 783 View FIGURES 782 – 795 , 805 View FIGURES 796 – 809 ); hind tibia with 8–9 inner spines; fastigium of vertex elongated ( Fig. 745 View FIGURES 743 – 759 )............................................................ 3

- Apical valves of penis large and stout, their ventral margins not turn to lateral sides, compressed antero-posteriorly along its length ( Fig. 89 View FIGURES 83 – 92 a); body small and clearly stouter ( Figs. 808–809 View FIGURES 796 – 809 ); hind tibia with 8 inner spines; fastigium of vertex short and wide ( Fig. 746 View FIGURES 743 – 759 )..................................................................( N. aserbus View in CoL sp. group)...15

3 Body relatively stouter and large ( Figs. 782–783, 786–795 View FIGURES 782 – 795 ); hind femur with a very distinct preapical notch, dorsal margin well developed, distinctly raised before the notch ( Fig. 747 View FIGURES 743 – 759 ); if preapical notch weak then body clearly larger; phallic complex large ( Figs. 82–85 View FIGURES 69 – 82 View FIGURES 83 – 92 ); hind tibia with 8–9 inner spines; mesosternal lobes in male not triangular ( Fig. 748 View FIGURES 743 – 759 )................................................................................................( N. serricollis View in CoL sp. group)...4

- Body very slender and relatively small; hind femur with a weak preapical notch, dorsal margin not raised before the notch ( Fig. 749 View FIGURES 743 – 759 ); phallic complex very small ( Figs. 86–88 View FIGURES 83 – 92 ); hind tibia with 9 inner spines; mesosternal lobes in male triangular ( Fig. 750 View FIGURES 743 – 759 )............................................................................... ( N. znojkoi View in CoL sp. group)...13

4 Hind femur with well developed dorsal margin, distinctly raised just before preapical notch which is very distinct and deep ( Fig. 751 View FIGURES 743 – 759 )...........................................................................................5

- Hind femur with low dorsal margin, preapical notch clearly weaker ( Fig. 752 View FIGURES 743 – 759 ).................................... 9

5 Frontal ridge distinctly incised just below the median ocellus ( Fig. 753 View FIGURES 743 – 759 ); fastigium of vertex more horizontal, slightly sloping; found in S. of Caucasia............................................................................... 6

- Frontal ridge slightly incised just below the median ocellus ( Fig. 754 View FIGURES 743 – 759 ); fastigium of vertex more sloping; found in N. of Caucasia................................................................................................ 7

6 Body stout and large; pronotal carinae blunt and low; slightly depressed between the median and lateral carinae ( Fig. 782 View FIGURES 782 – 795 ); dorsal margin of hind femur developed along the proximal part, not raised just before preapical notch ( Fig. 755 View FIGURES 743 – 759 ); pseudolophi on epiphallus large and very close each other, sometimes fused in the middle ( Fig. 82 View FIGURES 69 – 82 c); male phallic complex as in Fig. 82 View FIGURES 69 – 82 a, b, c; found in Armenia, Azerbaijan, Nakhichevan and E . Turkey.................................. N. serricollis View in CoL (F.-W.)

- Body slender and small; pronotal carinae very sharp and raised; strongly depressed between the median and lateral carinae ( Fig. 784 View FIGURES 782 – 795 ); dorsal margin of hind femur sharply developed, strongly raised just before preapical notch ( Fig. 751 View FIGURES 743 – 759 ); pseudolophi on epiphallus small and clearly seperated, never fused ( Fig. 83 View FIGURES 83 – 92 c); male phallic complex as in Fig. 83 View FIGURES 83 – 92 a, b, c; found in Georgia and Azerbaijan .................................................................. .. N. sanctidavidi (Shugurov) View in CoL

7 Hind femur short and broad, dorsal margin sharp, raised just before preapical notch ( Fig. 756 View FIGURES 743 – 759 ); median carina of pronotum in female clearly raised and convex, posterior end sharply projected in lateral view ( Fig. 791 View FIGURES 782 – 795 ); abdominal tergites more raised; inner and ventral surface of hind femur and hind tibia bright black............................. N. geniculatus Uvarov View in CoL

- Hind femur long and narrow, dorsal margin blunt, slightly raised just before preapical notch ( Fig. 757 View FIGURES 743 – 759 ); median carina of pronotum in female slightly convex or straighter, posterior end blunt, not projected in lateral view ( Figs. 787–788 View FIGURES 782 – 795 ); abdominal tergites less raised.................................................................................... 8

8 Female vertex strongly swollen, fastigium of vertex strongly depressed ( Fig. 758 View FIGURES 743 – 759 ); pronotum almost straight in lateral view ( Fig. 758 View FIGURES 743 – 759 ); inner and ventral surfaces of hind femur black, hind tibia blackish-grey.... N. daghestanicus loripes Mistshenko View in CoL

- Female vertex slightly swollen, fastigium of vertex slightly depressed ( Fig. 759 View FIGURES 743 – 759 ); pronotum convex in lateral view ( Fig. 759 View FIGURES 743 – 759 ); inner and ventral surfaces of hind femur black with red stains, hind tibia reddish-orange.................................................................................................. N. daghestanicus daghestanicus Uvarov View in CoL

9 Hind femur stout, proximal part wide, dorsal margin finely serrated and relatively sinuous especially in male, preapical notch more distinct ( Figs. 760–761 View FIGURES 760 – 781 )......................................................................... 10

- Hind femur slender, proximal part narrow and long, dorsal margin almost smooth and not sinuous, preapical notch quite weak ( Fig. 762 View FIGURES 760 – 781 )......................................................................................... 12

10 Fastigium of vertex acute in anterior part, sharply connected to frontal ridge; frontal ridge strongly projecting just above the median ocellus ( Figs. 763, 765 View FIGURES 760 – 781 ); eye in male large, its transversal diameter distinctly greater than vertex between the eyes and clearly wider than the distance between the frontal ridge and eye in lateral view ( Figs. 763, 765 View FIGURES 760 – 781 ); pronotal and abdominal carinae sharp; abdominal tergites with small posterior projection ( Figs. 794– 797 View FIGURES 782 – 795 View FIGURES 796 – 809 )................................... 11

- Fastigium of vertex less acute in anterior part, roundly connected to frontal ridge; frontal ridge not projecting just above the median ocellus ( Figs. 764, 766 View FIGURES 760 – 781 ); eye in male small, its transversal diameter smaller than vertex between the eyes and slightly wider than the distance between the frontal ridge and eye in lateral view ( Figs. 764, 766 View FIGURES 760 – 781 ); pronotal and abdominal carinae much blunter; abdominal tergites without posterior projection ( Figs. 792–793 View FIGURES 782 – 795 ) (in male, inner and ventral surfaces of hind femur reddish-black; hind tibia pale black in proximal part, turning to red in apical 1/3; tarsus red; in female, inner surface of hind femur pale black, turning to body color apically; ventral surface body color, its inner half slightly blackish and slightly reddish basally; hind tibia dark bluish-green with yellowish spines)........................ N. corrugatus Mistshenko View in CoL

11 Inner and ventral surfaces of hind femur black; hind tibia dark greyish-black in both sexes.................................................................................................... N. scabiosus scabiosus Mistshenko View in CoL

- Inner surfaces of hind femur black, its dorsal margin body color, inner half of ventral surface black in female, reddish in male, its outer half body color; hind tibia in male yellowish-cream turning to light red in apical half, in female proximal part blue turning to red in apical 1/4 including spines; tarsus red........................... N. scabiosus Mistshenkoi Descamps View in CoL

12 Body surface blackened without typical light band on abdomen in male ( Fig. 767 View FIGURES 760 – 781 ); inner and ventral surfaces of hind femur, hind tibia and tarsus completely black; pronotal and abdominal carinae and tubercles blunt ( Fig. 800–801 View FIGURES 796 – 809 ); frontal ridge wide, lateral margins blunt, not projecting forwards in lateral view; male phallic complex as in Fig. 85 View FIGURES 83 – 92 a, b, c N. pullus (Mistshenko) View in CoL

- Body of various shades of brown, with distinct typical light band on abdomen in male ( Fig. 768 View FIGURES 760 – 781 ); inner and ventral surfaces of

hind femur black with some reddish stains; hind tibia in male yellowish in basal half turning to orange-red in distal part, in female bluish in proximal part turning to red distally including spines; tarsus red in both sexes; pronotal and abdominal carinae and tubercles distinctly sharper ( Fig. 798 View FIGURES 796 – 809 ); frontal ridge narrow with sharp lateral margins, projecting farwards just above the median ocellus in lateral view; male phallic complex as in Fig. 84 View FIGURES 83 – 92 a, b, c.................... N. balachowskyi Descamps View in CoL 13 Median carina of pronotum with a distinct longitudinal sulcus ( Fig. 769 View FIGURES 760 – 781 ) (inner and ventral surfaces of hind femur black in both sexes; hind tibia pale black or reddish black turning to red at apex in male, blue or greenish turning to red at apex in female; tarsus red); male phallic complex as in Fig. 88 View FIGURES 83 – 92 a, b, c; found in the Elburs Mts., N. Iran.......... N. ebneri Ramme View in CoL

- Median carina of pronotum without longitudinal sulcus ( Fig. 770 View FIGURES 760 – 781 )............................................ 14

14 Inner and ventral surfaces of hind femur pale black or slightly reddish; hind tibia yellowish light orange in male, blue or greenish-blue with yellow spines in female; male phallic complex as in Fig. 86 View FIGURES 83 – 92 a, b, c...................... N. znojkoi Miram View in CoL

- Inner and ventral surfaces of hind femur black, outer half of ventral surface body color; inner surface of hind tibia pale black with dirty yellowish spines, dorsal surface dark greenish-blue or blackish; male phallic complex as in Fig. 87 View FIGURES 83 – 92 a, b, c........................................................................................... N. nanus Mistshenko View in CoL

15 Median carina of pronotum in female raised, clearly convex in lateral view ( Fig. 808 View FIGURES 796 – 809 ); mesosternal interspace narrower than mesosternal lobes ( Fig. 771 View FIGURES 760 – 781 ); inner and ventral surfaces of hind femur black; inner surface of hind tibia black, dorsal surface greyish-blue....................................................................... N. aserbus Mistshenko View in CoL

- Median carina of pronotum in female straighter in lateral view ( Fig. 811 View FIGURES 810 – 823 ); mesosternal interspace as wide as mesosternal lobes in female ( Fig. 772 View FIGURES 760 – 781 ); inner and ventral surfaces of hind femur pale black, ventral surface slightly reddish in female; inner surface of hind tibia black or strongly dark bluish in male, black turning to dark blue dorsally in female; male phallic complex as in Fig. 89 View FIGURES 83 – 92 a, b, c..................................................................... N. nodosus Mistshenko View in CoL

16 Dorsal margin of genicular lobe of hind femur raised, with sharp teeth ( Fig. 773 View FIGURES 760 – 781 ) (inner and ventral surfaces of hind femur black; inner surface of hind tibia black, dorsal surface grey like the is body color). Male unknown... N. crispus Mistshenko View in CoL

- Dorsal margin of genicular lobe of hind femur low, without sharp teeth ( Fig. 774 View FIGURES 760 – 781 )................................ 17

17 Arch of zygoma elongated, extending far beyond the posterior lobes of zygoma; apex of penis reaching to the level of arch of zygoma, its preapical part strongly widened and touching to the arch of zygoma ( Fig. 776 View FIGURES 760 – 781 ) (inner and ventral surfaces of hind femur and tibia black)............................................................................... 18

- Arch of zygoma very short and wide; apex of penis not reaching to the level of arch of zygoma, its preapical part slightly widened and not touching to the arch of zygoma ( Fig. 775 View FIGURES 760 – 781 ) (inner and ventral surfaces of hind femur and tibia black)............................................................................................. N. urmianus Ramme View in CoL

18 Female body small, compressed laterally; pronotum between the median and lateral carinae strongly sloping ( Fig. 777 View FIGURES 760 – 781 ); preapical notch of hind femur very shallow (see Ramme, 1951: 291, Abb. 80, fig. 4), inner surface of hind femur blackish-grey, ventral surface dirty yellowish-grey; hind tibia and tarsus blackish-grey ( Ramme, 1951). Male unknown.................................................................................................. N. keredjensis (Werner) View in CoL

- Female body large, depressed dorso-ventrally; pronotum between the median and lateral carinae depressed, slightly sloping ( Fig. 778 View FIGURES 760 – 781 ); preapical notch of hind femur much more distinct ( Fig. 780 View FIGURES 760 – 781 )........................................ 19

19 Arch of zygoma narrow, slightly raised and U-shaped basally ( Fig. 90 View FIGURES 83 – 92 a, b); anterior margin of pronotum in female strongly narrowed ( Fig. 779 View FIGURES 760 – 781 ); female hind femur as in Fig. 780 View FIGURES 760 – 781 ; male phallic complex as in Fig. 90 View FIGURES 83 – 92 a, b, c (inner and ventral surfaces of hind femur black; hind tibia black in male, black or blue turning to blue or greenish dorsally, rarely pink inner side of dorsal surface)............................................................................ N. iranicus (Werner) View in CoL

- Arch of zygoma not raised and V-shaped basally ( Fig. 776 View FIGURES 760 – 781 ); anterior margin of pronotum in female slightly narrowed ( Fig. 781 View FIGURES 760 – 781 ); female hind femur as in Fig. 774 View FIGURES 760 – 781 (inner and ventral surfaces of hind femur black; hind tibia dark bluish-green)....................................................................................... N. humerosus Mistshenko View in CoL

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Orthoptera

Family

Pamphagidae

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