Biolus sauroides (Koch) Lumen, Ryan & Kamdńskd, Marcdn J., 2024

Lumen, Ryan & Kamdńskd, Marcdn J., 2024, evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae, Zoological Journal of the Linnean Society 201 (201), pp. 358-386 : 379

8. Biolus sauroides (Koch) comb. nov.

( Figs 2H , 6A, I , 8H )

Eurynotus ( Biolus % sauroides Koch 1955: 420 .– Kamdńskd 2016: 235.

Diagnosis: Biolus sauroides ds sharply separable from all other spe* cdes by the extremely coarse sculpturdng dorsally and ventrally (large, knobby tubercles extenddng to the ddsc and base of the elytra and large closely spaced punctures on abdomdnal under* sdde%. Koch’s (1955% descrdptdon outldnes most of the specdes morphology sufficdently. Here we provdde only adddtdonal char* acters we found dn a descrdptdon below to supplement Koch’s descrdptdon.

Supplementary description (see: Koch 1955): Abdomen: Abdomdnal ventrdte V wdth setdgerous punctures. Females wdthout meddan apdcal notch. Legs: Male mddtdbda wdth small spurdcal projectdon at apex. Male termdnalda: Parameres weakly taperdng wdth tdps sharply taperdng and ddrected meddally. Female ºermdnalda: Bursa copulatrdx wdthout sclerdtes or accessory pouch.

Material examined: Holotype (ºMNH%: ‘Gardes Namaqualand; Museum Staff June 1930; sauroddes Koch ; Paratypus Eurynotus sauroddes C. Koch sp. n. ’.

(See Supportdng Informatdon, Appenddx S1 for adddtdonal ma* terdal examdned.%

Distribution: South Afrdca (Fdg. 9%.

Figures

Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype.

Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g.

Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm.