Tylodinus triumforium Luna-Cozar, 2014

Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, Zootaxa 3788 (1), pp. 1-63 : 32

16. Tylodinus triumforium Luna-Cozar , sp. nov.

( Figures 23, 26 , 56–57 , 109 , 130 )

Diagnosis. Length male, 3.9–4.5 mm, female, 3.5–4.2 mm. Width male, 2.3– 2.1 mm, female 1.9–2.1 mm. Body approximately1.9x longer than wide. Rostrum with small and shallow punctures. Elytra clothed with dark brown to light brown scales on disc, with yellow scales intermixed, denser on sides and in patches on intervals 1–3 at declivity, covering apical part of tubercle on I3 on middle of elytra and I4 on base of declivity, and covering tubercle on I2 almost completely. Male abdomen with ventrite 2 longer than 3–4 combined. Middle and hind tibiae with external tooth. Male genitalia with apical process in ventral view small ( Figure 109 ). Female genitalia with median membranous area of sternite eight as long as basal plate, spermatheca shallowly curved.

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1500–2200 m.

Derivation of specific name. Name derived from the Latin word triumphus = triumph, and named after the El Triunfo Biosphere Reserve, the protected natural area where the species was collected.

Material examined. Total 11 males, 13 females. Holotype male ( CMNC): México: Chiapas, Mpio Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal , elev. 2220 m (15.70819°N 92.93070°W), 21.v.2008, coll. R . S. Anderson, mossy oak forest litter. Paratypes: México: Chiapas, Angel Albino Corzo, Reserva Biosfera El Triunfo, Campamento El Quetzal , elev. 2220 m (15.70819°N 92.93070°W), 21.v.2008, coll. R . S. Anderson — 1 female ( CMNC); La Concordia , 4 km SE Custepec, elev. 2125 m (15.70658°N, 92.93126°W), 20.v.2008, coll. LLAMA (Wa-A-03-2-29) — 1 male ( CMNC); La Concordia , 2 km SE Custepec, elev. 1520 m (15°43'13.58''N, 92°57'4.18''W), 17.v.2008, coll. LLAMA (Wa-A-02-1-39) — 2 males, 1 female ( CMNC, ECOS); same locality (15.72051°N, 92.95164°W), LLAMA (Wa-A-02-1-42) — 1 male, 1 female ( CMNC, BMNH); same locality, elev. 1700 m (15.71595°N, 92.93842°W), LLAMA (Wa-A-02-2-03) — 1 female ( CMNC); same locality (15.71603°N 92.93808°W) LLAMA (Wa-A-02-2-19) — 1 female ( ECOS); same locality (15.71580°N 92.93811°W), LLAMA (Wa-A-02-2-24) — 1 male ( ECOS); same locality (15.71576°N 92.93812°W), LLAMA (Wa-A-02-2-25) — 1 female ( CMNC); same locality, elev. 2125 m (15.70708°N, 92.93131°W), LLAMA (Wa-A- 03-2-17) — 1 male ( CMNC); same locality, elev. 2140 m (15.70925°N, 92.92929°W), LLAMA (Wa-A-03-1-48) — 1 female ( CWOB); 3.5 km ESE Custepec, Campamento El Quetzal, elev. 1850 m (15°43'.1''N, 92°55'59.9''W), 17.vii.2007, coll. M. G. Branstetter — 1 male, 1 female ( CMNC); Reserva de la Biósfera El Triunfo, Campamento El Quetzal , elev. 1830 m (15°43'17''N, 92°56'20.8''W), 20.v.2008, coll. R . S. Anderson — 1 female ( CMNC); same locality, 22.v.2008, coll. R . S. Anderson — 1 female ( CMNC); same locality, elev. 1530 m, 18.vii.2007, coll. R . S. Anderson — 1 male ( CMNC); same locality, elev. 1800 m, 18.vii.2007, coll. R . S. Anderson — 1 female ( CMNC); same locality (15.72188°N, 92.93677°W), elev. 1900 m, 19.v.2008, coll. R . S. Anderson — 1 male ( CMNC); same locality (15.72178°N, 92.94544°W), elev. 1700 m, 17.v.2008, coll. R . S. Anderson — 2 females ( CMNC, ECOS); same locality, elev. 2177 m, 23.v.2008, coll. J. L. Cozar — 1 male ( CMNC) .

Figures

FIGURES 20–27. 20–21. Tylodinus spiniventris: 20. meso- and metathorax (male) 21. ventrite 5 (female), 22. Tylodinus elongatus ventrite 5 (female), 23. Tylodinus triumforium, ventrite 2 (male). 24. Tylodinus complicatus, ventrite 2 (female). 25. Tylodinus complicatus, ventrite 2 (male), 26. Hind tibia of Tylodinus triumforium, 27. Front tibia of Tylodinus rugosus.

FIGURES 52–59. Dorsal and lateral habitus: 52–53. Tylodinus coapillensis, 54–55. Tylodinus leoncortesi, 56–57. Tylodinus triumforium, 58–59. Tylodinus sepulturaensis.

FIGURES 100–109. Ventral and lateral habitus of median lobe: 100. Tylodinus andersoni, 101. Tylodinus cavicrus, 102. Tylodinus pseudocavicrus, 103. Tylodinus rugosus, 104. Tylodinus exiguus, 105. Tylodinus ixchel, 106. Tylodinus jonesi, 107. Tylodinus variabilis, 108). Tylodinus leoncortesi, 109. Tylodinus triumforium.

FIGURES 130–131. Map showing collecting sites. 130. Occurrences in México (Chiapas) and Central America of: Tylodinus zilchi (El Salvador, Honduras); Tylodinus andersoni (México), Tylodinus canaliculatus (México, Guatemala), Tylodinus triumforium (México), Tylodinus sepulturaensis (México), Tylodinus cavicrus (Guatemala), Tylodinus pseudocavicrus (México); 131. Occurrences in Chiapas, México of: Tylodinus complicatus, Tylodinus ixchel, Tylodinus dominicus, Tylodinus immundus, Tylodinus porvenirensis, Tylodinus branstetteri.