Leptotarsus ibericus, Ribeiro, Guilherme C. & Lukashevich, Elena D., 2014

Ribeiro, Guilherme C. & Lukashevich, Elena D., 2014, New Leptotarsus from the Early Cretaceous of Brazil and Spain: the oldest members of the family Tipulidae (Diptera), Zootaxa 3753 (4), pp. 347-363 : 357

Leptotarsus ibericus sp. nov.

( Figs 7, 8 , 22–25 )

Type material. Holotype: LH 28206± (part and counterpart of entire male specimen; preserved structures include the long antenna and terminalia). Paratypes: LH 29995± (part and counterpart of isolated wing) and LH 18122± (part and counterpart of adult). East-central Spain, Las Hoyas; La Huérguina Limestone Formation, Late Barremian, Early Cretaceous. Housed in Museo de Ciencias de Castillia–La Mancha.

Etymology. Named after Iberian Mountain Range.

Diagnosis. This species differs from species of Leptotarsus from the Cretaceous of Brazil by absence of dark costal field, and from L. buscalioniae sp. nov. by the shorter distal section of M1+2 and distal sections of veins M3 and M4 almost parallel.

Description. Head. Head capsule with an anterodorsal protuberance; antennae long and slender, bearing long and slender flagellomeres (total number of flagellomeres unknown); scape stout and long, swollen in middle, 3x longer than pedicel; vestiture of flagellar segments long, delicate and abundant, without visible verticils, length of macrosetae ca. 4.0x the diameter of flagellomere (visible on short portion of antenna); nasus distinct, rostrum ca. 0.5x the total length of head. Thorax stout, as long as wide; transverse suture distinct; praescutal longitudinal stripes clearly delimited. Wing transparent except for dark pterostigma. Sc reaching wing margin at the level of first bifurcation of Rs and beyond level of the tip of Cu; sc-r present, positioned near the tip of Sc; Rs subequal to R2+3 stem; R1 reaching wing margin distal to the level of R2+3 bifurcation; r-r oblique, linking R1 to R2, R2 oblique; medial vein four-branched; discal cell wide; distal sections of veins M3 and M4 subparallel, subequal; long m-cu linking Cu to the base of M4. Abdomen stout, ca. 1.5x longer than thorax; gonocoxite probably conical, as long as wide.

Measurements. Antenna length, 19.4. Body length, 7.8. Wing length/width, 6.9–7.9/2.1–2.5.

Figures

FIGURES 5 – 9. Wing venation of Leptotarsus spp. from Las Hoyas, Early Cretaceous, Spain. 5 — L. buscalioniae sp. nov., holotype LH 35088, right wing. 6 — L. buscalioniae sp. nov., holotype LH 35088, left wing. 7 — L. ibericus sp. nov., holotype LH 28206. 8 — L. ibericus sp. nov., paratype LH 29995. 9 — L. contractus sp. nov., holotype LH 15419. Scale bar 1 mm.

FIGURES 22 – 25. Leptotarsus ibericus sp. nov., from Las Hoyas, Early Cretaceous, Spain: 22 — Holotype LH 28206, total view (antenna truncated). 23 — Holotype LH 28206, detail of head. 24 — Holotype LH 28206, head with antennae. 25 — Paratype LH 29995. Abbreviations: comp eye: compound eye; na: nasus; plp: palpus; scp: scape.