Mycomya paguma , Väisänen, Rauno, 2013

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 2. Subgenera Calomycomya, Cymomya, Neomycomya and Pavomya subg. n., Zootaxa 3666 (3), pp. 301-318: 311-312

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Mycomya paguma

sp. n.

Mycomya paguma  sp. n.

Figs. 6View FIGURE 6 A –C

Material examined. Holotype. 3. NEPAL, 27 ° 56 'N, 85 °00'E, Mal. tr. 9, 9900 ', 28.V. 1967, Can. Nepal Exp. (in CNC). Paratypes (3 only). 28 °00'N, 85 °00'E, Mal. tr. 7, 9900 ', 26.V. 1967, Can. Nepal Exp., 1 3 (in MZH); Gupa Pokali, 27 ° 17 'N, 87 ° 30 'E, 2900 m—Chouki. 27 ° 12 'N, 87 ° 28 'E, 2700 m, 16.VI. 1972, J. Emoto, 1 3 and 1 Ƥ (in KUC).

Description. 3. Head. Palp and other mouthparts yellow, clypeus and face brownish, posterior parts of head brown. Antenna brown, scape, pedicel and base of 1 st flagellomere yellow. Length of 1 st flagellomere about 2.5 x, 2 nd flagellomere 2 x, its width. Thorax. Pronotum yellow, with 3 long setae. Scutum brown, with anterolateral corners yellowish. Anepisternum brownish. Preepisternum brownish. Scutellum brownish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 3.6–4.1 mm. Wing hyaline. Sc ending in R 1 near middle of small cell, Sc 1 missing. Apical part of Sc bearing 4–10 small setae. Small cell 1.5–2 times as long as wide. Cu fork slightly proximal to M fork. M ratios: 1.00, 1.38. Cu ratios: 0.66, 1.06. Small setae: M petiole: 0; M 1: +; M 2: 5; Cu petiole: 0; Cu 1: 2–7; Cu 2: 2–8. Halter pale yellowish. Legs. Coxae yellow, femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt 1:t 1 = 0.80–0.85, bt 2:t 2 = 0.65–0.73, bt 3:t 3 = 0.58– 0.61. Abdomen. Tergites brown, 3–5 with small yellow anterolateral spots, sternites 1–5 yellow with brownish to brown posterior margins, 6–7 brown. Hypopygium. Figs. 6View FIGURE 6 A –C, yellow. Sternal lateral appendage short, apically rounded, widest basally ( Fig. 6View FIGURE 6 B). Sternal submedian filament almost straight, distinctly longer than sternal lateral appendage ( Figs. 6View FIGURE 6 A –B). Apex of aedeagus rounded in lateral view ( Fig. 6View FIGURE 6 C). Female. Wing length 4.5 mm. Pronotum yellow. Scutum brown with 2 yellowish longitudinal stripes. Anepisternum light brownish. Preepisternum yellowish, ventral half brownish. Scutellum, laterotergite and mediotergite brownish to brown. Abdominal tergites 1–5 brown with yellowish anterolateral spots, 6–7 brown, sternites lighter. Leg ratios: bt 1:t 1 = 0.73, bt 2:t 2 = 0.68, bt 3:t 3 = 0.58.

Discussion. Mycomya paguma  differs from the rest of the Himalayan species in having wide, apically rounded sternal lateral appendages ( Fig. 6View FIGURE 6 B) of the shape of a reversed U (a bit similar to that of the much larger M. panthera  ), and the almost straight sternal submedian filaments, which are distinctly longer than the sternal lateral appendages ( Figs. 6View FIGURE 6 A –B). The male hypopygium of M. paguma  resembles that of the Nearctic M. obliqua (Say, 1824)  , but differs in the shape of the sternal submedian filaments, which are curved in M. obliqua  . Mycomya paguma  seems to be closely related to M. gutianshana  Wu & Yang, which was described from Zhejiang Province, eastern China, at the altitude of 400–500 m (Wu & Yang 1994). Both species have long and almost straight sternal submedian filaments and the general appearance of the hypopygium is quite similar, but the sternal lateral appendage of M. gutianshana  is said to be slender (unfortunately the hypopygium was not figured in a lateral view), while it is wide and rounded in M. paguma  . Mycomya gutianshana  is smaller (wing length 3.1–3.8 mm) than M. paguma  (3.6–4.1 mm), and its small cell is shorter (1.0– 1.5 x as long as wide). Although the differences between are M. paguma  and M. gutianshana  are not great, they are considered to represent two separate species also taking into account that M. paguma  is a high altitude species and their known distributions seem to be widely disjunct.

Etymology. The species name is a noun in apposition and refers to the Himalayan palm civet (genus Paguma  ).