Mycomya

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 2. Subgenera Calomycomya, Cymomya, Neomycomya and Pavomya subg. n., Zootaxa 3666 (3), pp. 301-318 : 303

Key to the subgenera of Mycomya and Himalayan species of subgenera Calomycomya , Cymomya , Neomycomya and Pavomya

1. Coxa 2 with a spur (see Väisänen 1984: figs. 377, 436, 445, 597, 825).......................................... 2

- Coxa 2 without a spur............................................................................... 5

2. (1) Abdominal tergites yellow with dark posterior margins; abdominal tergite 8 bare; tergal part of hypopygium with a fork-like median structure and a specialized inner structure including a pair of strong spurs ( Figs. 2 A, and 1A, 2C, 3A; Väisänen 1984: e.g. figs. 833, 842, 851); ocellar prominence usually distinctly darker than posterior parts of head........................................................................................... (subgenus Calomycomya ) 3

- Abdominal tergites entirely dark or light, or dark with paler posterior margins; abdominal tergite 8 with setae; tergal part of hypopygium without such structures; ocellar prominence not distinctly darker than posterior parts of head................................................................................ subgenus Mycomya (see Väisänen 1996)

3. (2) Tergal fork of hypopygium basally wide, U-shaped, apically 1.2x as wide as its length ( Fig. 2 A)......... M. ratufa sp. n.

- Tergal fork of hypopygium basally narrow, V-shaped, apically narrower than its length ( Figs. 1 A, 3A)............... 4

4. (3) Both branches of gonostylus relatively long and slender ( Fig. 3 D); aedeagus with two short bilobed spurs ( Fig. 3 E).............................................................................................. M. wah sp. n.

- Slender branch of gonostylus much shorter than the broader one ( Fig. 1 C); aedeagus with two long spurs ( Fig. 1 D).......................................................................................... M. marmota sp. n.

5. (1) Mediotergites with small setae or bare; coxa 1 with or without a dense patch of short, seldom longer, setae on anteromedial surface (Väisänen 1984: figs. 104, 435); tergal part of hypopygium usually with 2 groups of dark cones, often with lateral sabre-like spines (e.g. Väisänen 1984: figs. 76, 119, 254); sternal synsclerite not medially deeply emarginated, with setose lobe-like submedian appendages (e.g. Väisänen 1984: figs. 133–136)...................... subgenus Mycomya (p.p.)

- Mediotergite bare; coxa 1 without specialized setae; tergal part of hypopygium without such cones or spines; sternal synsclerite deeply emarginated or divided into two parts, without such appendages.................................... 6

6. (5) Abdominal tergite 8 bare ( Fig. 10 C), sternite 8 with setae ( Fig. 10 D).............................................................................................. ..subgenus Pavomya subg. n.: Mycomya kambaitiensis sp. n.

- Abdominal tergite 8 with long apical setae (Väisänen 1984: figs. 896, 919, 974), sternite 8 usually bare (with setae only in subgenus Myacymo)................................................................................. 7

7. (6) Sternal synsclerite with long wide lateral appendages (e.g. Fig. 5 C); tergal part of male hypopygium without lateral appendages............................................................................................. 8

- Sternal synsclerite without large lateral appendages; tergal part of hypopygium with lateral appendages (e.g. Väisänen 1984: figs. 964–965, 1059–1060, 1066)..................................................................... 13

8. (7) Tergal part of hypopygium without a comb, with groups of strong submedian and lateral spines (e.g. Väisänen 1984: figs. 914, 917, 926, 928, 937–939, 943); sternal lateral appendages with strong spines (e.g. Väisänen 1984: figs. 916, 921, 927, 934, 942).................................................. subgenus Coheromyia (Holarctic and Neotropical)

- Tergal part of hypopygium with an apicomedial comb of setae, without groups of strong spines (Väisänen 1984: figs. 911– 913); sternal lateral appendages without strong spines..................................................... 9

9. (8) Setae of tergal comb strong and very long, almost as long as width of hypopygium (Väisänen 1984: figs. 911–913)................................................................................subgenus Tlingitmyia (Nearctic)

- Setae of tergal comb short, spine-like (e.g. Väisänen 1984: fig. 892, 901).................... (subgenus Cymomya ) 10

10. (9) Sternal lateral appendage long, narrow and its apical part curved ventrally ( Fig. 5 C).................... M. cuon sp. n.

- Sternal lateral appendage comparatively short, wide and straight, apically rounded ( Figs. 4 B, 6B, 7B)............... 11

11. (10) Sternal submedian filament distinctly (about 1.2x) longer than sternal lateral appendage ( Fig. 6 A)....... M. paguma sp. n.

- Sternal submedian filament not longer than sternal lateral appendage ( Figs. 4 A–B, 7 A–B)....................... 12

12. (11) Sternal lateral appendage widest at midlength ( Fig. 4 B); wing length 3.2–4.1 mm .................... M. aonyx sp. n.

- Sternal lateral appendage widest basally ( Fig. 7 B); wing length 4.6 mm .......................... M. panthera sp. n.

13. (7) Tergal part of hypopygium without tergal combs......................................................... 14

- Apical part of tergal part of hypopygium with at least one dense tergal comb of short spines (e.g. Väisänen 1984: figs. 1042, 1045, 1064)...................................................................................... 15

14. (13) Tergal part of hypopygium with relatively small submedian lobes, lateral appendages long, well-developed, with specialized setae (Väisänen 1984: figs. 946, 948, 949, 951); abdominal sternite 8 with setae......... subgenus Myacymo (Nearctic)

- Tergal part of hypopygium with large submedian lobes, lateral appendages small ( Figs. 8 A, 8C); abdominal sternite 8 bare............................................................. subgenus Neomycomya : M. fimbriata (Meigen)

15. (13) Tergal part of hypopygium with two dense submedial, apical tergal combs (e.g. Väisänen 1984: figs. 985, 989, 1042, 1045)................................................................................ subgenus Mycomyopsis

- Tergal part of hypopygium with a dense wide apical tergal comb (e.g. Väisänen 1984: figs. 1064, 1071, 1075).............................................................................................. subgenus Lycomya

Figures

FIGURE 1. Mycomya marmota sp. n., male hypopygium. A – B tergal and sternal views; C gonostylus, twice enlarged; D apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.

FIGURE 2. Mycomya ratufa sp. n., male hypopygium. A – C tergal, sternal and lateral views, tf—tergal fork; D apex of sternal lateral appendage, twice enlarged; E gonostylus, twice enlarged; F apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.

FIGURE 3. Mycomya wah sp. n., male hypopygium. A – C tergal, sternal and lateral views; D gonostylus, twice enlarged; E apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.

FIGURE 4. Mycomya aonyx sp. n., male hypopygium. A – B sternal and lateral views; C apex of apex of aedeagus, lateral view, twice enlarged; D tergite 8. Scale 0.2 mm.

FIGURE 5. Mycomya cuon sp. n., male hypopygium. A – C tergal, sternal and lateral views, sla—sternal lateral appendage; D apex of aedeagus, lateral view, twice enlarged; E tergite 8. Scale 0.2 mm.

FIGURE 6. Mycomya paguma sp. n., male hypopygium. A – B sternal and lateral views, ssf—sternal submedian filament; C apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.

FIGURE 7. Mycomya panthera sp. n., male hypopygium. A – B sternal and lateral views, C apex of aedeagus, twice enlarged, D tergite 8. Scale 0.2 mm.

FIGURE 8. Mycomya fimbriata (Meigen, 1818), male hypopygium. A – C tergal, sternal and lateral views; D – E gonostylus in different positions; F apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.

FIGURE 10. Mycomya kambaitiensis sp. n., male hypopygium. A lateral views; B gonostylus, twice enlarged; C tergite 8; D sternite 8. Scale 0.2 mm.