Daedalma parvomaculata Krüger

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 22

Daedalma parvomaculata Krüger

( Figs. 4A, 4B, 10F, 14A, 23A)

Daedalma parvomaculata Krüger, 1924: 47 .

Daedalma dinias parvomaculata Krüger ; Adams, 1986: 253; d’Abrera, 1988: 844; Salazar, 1993: 39.

Daedalma dinias form parvomaculata Krüger, 1924: 47 .

Daedalma parvomaculata Krüger ; Pyrcz, 1999: 228; Pyrcz & Wojtusiak, 1999: 206; Prieto, 2003: 209, fig.1; Pyrcz, 2004: fig. 2; Vargas & Salazar, 2004: 253.

Remarks: D. parvomaculata is currently considered monotypic. It most closely resembles D. eliza in wing shape and HWV colour pattern, but is immediately recognised by the brighter orange FW patch, which is dark brick red in D. eliza , and by significant differences in the female (see under D. eliza ).

Material examined: COLOMBIA: LECTOTYPE male: Cauca, Western Cordillera, West slopes, La Paz, 2000 m, 22.IX.1917, E. Krüger leg. , MIIZ (Designated by Pyrcz , 1999: 228); PARALECTOTYPE male: same data but 21.IX.1917 ; 1 female: same data but 20.IX.1917 ; 1 male: Cauca, Reserva Forestal Tambito , 2050 m, 14.II.1997, T. Pyrcz leg. , MZUJ; 1 male: same data but 2400 m, 04.II.1997 ; 1 male: same data but 1900 m, 09.II.1997 ; 1 male: same data but 1950 m, 09.II.1997 ; 1 male: same data but 2000 m, 09.II.1997 ; 1 male: same data but 2250 m, 09.II.1997 ; 1 male: same data but 2300 m, 09.II.1997 ; 1 male: same data but 1950 m, 10.II.1997 ; 1 male: same data but 2000 m, 10.II.1997 ; 1 male: same data but 2300 m, 10.II.1997 ; 1 male: same data but 2000 m, 12.II.1997 ; 1 male: same data but 2150 m, 12.II.1997 ; 1 male: same data but 2300 m, 12.II.1997 ; 1 male: same data but 2400 m, 12.II.1997 ; 2 males: same data but 2300 m, 13.II.1997 ; 1 male: same data but 2000 m, 14.II.1997 ; 1 male: same data but 2150 m, 14.II.1997 ; 1 male: same data but 2250 m, 14.II.1997 ; 1 male: same data but 1950 m, 01.III.1997 ; 1 male: same data but 2000 m, 01.III.1997 ; 1 male: same data but 2050 m, 01.III.1997 ; 2 males: same data but 2300 m, 01.III.1997 ; 1 male: same data but 2400 m, 01.III.1997 ; 1 male: same data but 2000 m, 02.III.1997 ; 1 male: same data but 2300 m, 02.III.1997 ; 1 male: same data but 1900 m, 03.III.1997 ; 1 male: same data but 1950 m, 03.III.1997 ; 1 male: same data but 2400 m, 03.III.1997 ; 1 male: same data but 1950 m, 04.III.1997 ; 1 male: same data but 1950 m, 05.III.1997 ; 2 males: same locality but no other data, MZUJ ; 1 male: Cauca, P. N. Munchique, La Romelia , 2800 m, 20.I.2001, C. Prieto leg. ; 1 male: same data but 09.VI.2006 ; 2 males: Cauca, P. N. Munchique, Veinte de Julio , 2700 m, 15.V.2006, C. Prieto leg. ; 1 female: Valle del Cauca, Queremal, Cerro Tokio , 3º 28' 00 N, 76º 42' 00 W, 2000 m, 25.X. 2008, C. Prieto leg. , CPC.

Redescription: MALE ( Fig. 4A): Head: Eyes dark brown; palpi rufous brown; antennae dorsally medium brown, ventrally orange brown. Thorax: blackish brown; legs medium brown. Abdomen: dorsally blackish brown, laterally and ventrally slightly lighter and duller. Wings: FW length 25–27 mm, mean: 25.8 mm, n=39; FWD blackish brown; oval orange patch at base of space M3-Cu1 entering marginally discal cell and space Cu1-C2. HWD uniform blackish brown. FWV: steely brown, lustrous; orange patch slightly smaller than on upperside; subapical and apical area chocolate brown and with short, irregular milky white lines; three minute blackish brown spots in R4-R5, R5-M1 and M1-M2. HWV: predominantly crimson brown with some orange speckling, silvery and postbasal, postmedian and submarginal irregular and discontinuous lines; a row of dark brown ocelli in postmedian to submarginal area. Genitalia ( Fig. 10F): Characterised by a particularly stout uncus, a feature which strongly suggests an affinity with D. eliza . Valvae are shortened in apical half and terminated with a prominent tip pointed upwards, ampulla has a slightly serrate surface.

FEMALE ( Fig. 4B): Differs from male in lighter, medium brown FW and HWD, and in having orange FWD patch twice as large and equally conspicuous on ventral surface (faint and suffused with brown on male FWV); HWV colour pattern much lighter than in male. Genitalia ( Fig. 14A): Sinus vaginalis small. Marginal parts of Vshaped batten on lamella postvaginalis constitute a heavily sclerotized, very characteristic trapezoidal frame. Papillae anales each with small flattened lobe on ventral part and a bump on outer surface. Apophyses posteriores as short triangular plates. Ductus bursae four times longer than antrum, with delicate parallel wrinkles, attached to corpus bursae asymmetrically. Colliculum well developed, wall of lamella postvaginalis posterior to opening of antrum with few wrinkles. Hump in middle of lamella postvaginalis slightly concave when viewed ventrally. Bursa copulatrix with two ribbon-like, parallel signa consisting of minute teeth.

Remarks: Daedalma parvomaculata is a species endemic to the Colombian Chocó, and is known to occur in the departments of Cauca, Valle del Cauca, Chocó, and Risaralda ( Vargas & Salazar, 2004). Described originally as a form of D. dinias , it was considered by Adams (1986) as a subspecies of D. dinias . Daedalma parvomaculata and D. dinias are, in fact, widely parapatric on the Pacific slopes of the Colombian Western Cordillera, the latter occurring at higher elevations, and D. parvomaculata was therefore treated as a valid species by Pyrcz (1999). It is apparently most closely related to D. eliza , from which it is separated geographically by the Río Patia Valley. Daedalma parvomaculata occurs in low to mid-elevation perhumid cloud forests between 1800–2200 m ( Pyrcz & Wojtusiak, 1999). Sexual dimorphism appears less pronounced than in other species of the “ dinias group", with the female differing from the male only in having a wider FW orange patch and lighter HWV.