Daedalma, Hewitson, 1858

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 23-24

Daedalma vertex vertex Pyrcz

( Figs. 3E, 3F, 10A, 15D)

Material examined: PERU: HOLOTYPE male: Amazonas , Molinopampa-Granada , 3065 m, 03.VII.1998, T. Pyrcz & J. Wojtusiak leg., MUSM; 1 male: Amazonas, Molinopampa-Granada, 2915 m, 29.VIII.1998, T. Pyrcz & J. Wojtusiak leg., TWP; 1 male: same data but 2800–3100 m, 20.VIII.1998, TWP; 1 male: same data but 3000– 3050 m, B. Calderón leg., TWP; 17 males: same data but 3100–3250 m, M. Tafur leg., 10 TWP, 7 MUSM; 1 male: Peña Blanca, Río Chido , 2800 m, 29.VI.1999, B. Calderón leg., TWP; 3 males: Road Leimebamba-Balsas, 3550– 3650 m, 1–15.XII.2001, B. Calderón leg., MBLI; 1 male: Molinopampa-Granada trail, 2500 m, X.2000, B. Calderón leg., MB; 1 male: same data but 3050 m, MBLI; 4 males: same data but 3150–3250 m, MBLI; 1 male: same data but 3400 m, MBLI; 1 male: Molinopampa 2900–3000 m, IX.2000, B. Calderón leg., PBF; 1 male: Pomacochas, La Sonada, IX–X.2000, B. Calderón leg., PBF; 1 female: Molinopampa-Granada, 3100–3250 m, III.2003, M. Tafur leg., TWP; 1 female: Peña Blanca, Río Chido, 2500–2900 m, VI.2002, B. Calderón leg., TWP; 1 female: Peña Blanca, Laguna de Pomacochas, 2900–3050 m, VI.2000, B. Calderón leg., MBLI; 1 female: Pueblo María, Luya, 3200 m, XI.1998, B. Calderón leg., MBLI; 1 female: Molinopampa-Granada trail, 3150–3250 m, X. 2000 m, B. Calderón, MBLI; 1 female: same data, PBF; 1 female: Molinopampa, 2900–3100 m, I.2001, B. Calderón leg., PBF; 1 female: Pomacochas, 3200 m, 13.VI.2000, B. Calderón leg., PBF; 1 female: Amazonas, Molinopampa- Granada, 3100–3250 m, III.2003, M. Tafur leg., TWP (all PARATYPES); 1 female: La Sonada, Pomacochas, IX– X.2000, B. Calderón leg., PBF; 1 female: Molinopampa, 2900–3100 m, I.2001, B. Calderón leg., PBF; 1 female: Pomacochas, 3200 m, 13.VI.2000, B. Calderón leg., PBF.

Redescription: MALE ( Fig. 3E): FWD dark chocolate-brown, lustrous; transverse oblong, median patch extending from near costa, across discal cell to mid-cell Cu1-Cu2, with a triangular incision at base of vein Cu1 and a dot enclosed within orange patch in cell Cu1-Cu2; faint, barely visible postdiscal orange line from vein M1 to M3; fringes brown and yellowish in interspaces. HWD uniform dark chocolate-brown, lustrous, except for a pale orange costal lightening before apex; fringes brown and yellowish in apical one-third. FWV dark chocolate-brown; orange elements reflected from upperside, but post-discal line slightly better marked; distally, apical brown area lighter and suffused with magenta and a fine brown submarginal line; two brown faint ocelli in cells R5-M1 and M1-M2. HWV pattern an extremely complex mosaic of black, brown and beige with traces of magenta and a series of triangular, shining yellow submarginal patches, similar to D. boliviana except that two submarginal shining yellow triangular patches are less elongated and slightly larger; postmedian line edged basally with black; ocellus in cell Cu1-Cu2 black, without white pupil. Genitalia ( Fig. 10A): Uncus moderately stout; gnathos stout in basal, thin in distal half; valvae slightly narrower in distal than in medial part, with a blunt apex and a sharp tip pointing upwards; dorsal surface smooth; saccus intermediate in length; aedeagus slightly contorted in distal one-third, similar in length to valva.

FEMALE ( Fig. 3F): FWD patch light orange. HWD orange patches extending from costa towards vein Cu 2 in postmedian to submarginal area, gradually darkening, becoming rusty orange, and fading away. Genitalia ( Fig. 15D): Sinus vaginalis wide. V-shaped batten on lamella postvaginalis wide and heavily sclerotized, with widely rounded central hump. Papillae anales trapezoidal in lateral view, ventrally separated by a narrow gap. Apophyses posteriores short, triangular and well sclerotized. Cuticular slat connecting eighth tergite with sinus vaginalis strongly sclerotized along its posterior edge. Bursa copulatrix small and elongated. Ductus bursae very short. Antrum short, its diameter larger than that of ductus bursae. Ductus seminalis very short, connecting with antrum posteriorly at colliculum near ostium. Long axes of bursa copulatrix and ductus bursae not aligned, bursa directed ventrally. Bursa copulatrix with two ribbon-like, parallel signa consisting of minute teeth.

Remarks: The nominate subspecies of D. vertex was described from the highlands of Chachapoyas in northern Peru, and later found in the area of Abra Barro Negro in the Central Cordillera, also in Amazonas. The latter individuals differ in having a slightly narrower FW subapical orange band, that is, however, not as narrow as in milleri n. ssp. Daedalma vertex is a high elevation species, usually found above 2600 m, and often above 3000 m ( Pyrcz, 2004).