Daedalma drusilla tinta Pyrcz, 2011

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 29-30

Daedalma drusilla tinta Pyrcz , n. ssp.

( Figs. 7G, 7H, 10E)

Material examined: COLOMBIA: HOLOTYPE male: Antioquia, San Felix , 2800–3000 m, 29.III.2003, G. Rodríguez leg., red, rectangular label saying : Holotype, MZUJ (to be deposited in MHN-UNC) ; PARATYPES (10 males and 1 female): 1 male: same data as the holotype, GRM ; 1 male: Antioquia, San Felix , 2800–3000 m, 29.III.2003, G. Rodríguez leg., TWP ; 1 male: Antioquia, San Felix , 2900–3100 m, 07.VIII.2000, G. Rodríguez leg., GRM ; 1 male: Antioquia, San Felix , 2900–3100 m, 07.VIII.2000, G. Rodríguez leg., TWP, (prep. genit. TWP-03 / 28.05.2008) ; 6 males: same data, MBLI ; 1 female: Antioquia, San Felix , 2900 m, 10.VIII.2000, G. Rodríguez leg., GRM .

Description: MALE ( Fig. 7G): Head, thorax and abdomen: not differing from nominate subspecies. Wings (FW length: 24–25.5 mm, mean: 25.2 mm, n=8): FWD and HWD uniform blackish brown, a shade darker than in nominate subspecies. FWV ground colour blackish brown, noticeably darker than chestnut of nominate; patches in distal part of discal cell and submarginal area fainter than in nominate, barely lighter than ground colour. HWV ground colour also darker than in nominate subspecies, chocolate brown; submarginal milky white patches slightly larger, particularly one in M2-M3 more elongated. Genitalia ( Fig. 10E): Uncus stout, longer than dorsum of tegumen, slightly thicker than in nominate subspecies; gnathos 2/3 length of uncus, thin, curving upwards; saccus deep, about same length as uncus; valvae elongated, wider than in nominate subspecies, with a smooth ampulla and a prominent apical hook curved upwards; aedeagus smooth, about same length as valvae, gently arched in middle.

FEMALE ( Fig. 7H): FW length: 26 mm; FWD ground colour dark brown; wine-red patches suffused with brown covering most of distal half: an irregular patch in distal corner of discal cell, an elongate subapical patch, and a large, roughly rectangular patch extending in postmedian to submarginal area between vein M3 and anal margin; apical area dusted with light brown scales. HWV ground colour medium brown, slightly lighter than on FW; reddish markings slightly lighter than on FW with an orange sheen, forming a diffuse mid discal cell patch and covering area between postdiscal and submarginal line, a series of five brown postdiscal patches, three of which in Rs- M1, M1-M2 and M2-M3 oval, two in M3-Cu1 and Cu1-Cu2 lunular and much larger. FWV colour pattern same as on dorsal surface, with a magenta patch on costa near apex, three minute blackish subapical ocelli, a chocolate brown area on distal margin between veins M1 and M2, turning into a yellowish distal margin gradually narrowing towards tornus. HWV colour pattern not differing noticeably from females of other subspecies. Genitalia: Not examined.

Etymology: The epithet of this subspecies derives from the noun tinto, which in Spanish means red wine, and refers to the diagnostic colour of the upperside patches of the female.

Remarks: This subspecies differs from other races of D. drusilla , mostly by the unusual wine-red markings on the dorsal surface of the female. Moreover, the blackish pattern in the male is darker than in other subspecies. All known individuals were collected on locally isolated mountain ridges, ascending to 3100 m around the city of Medellín, the capital of Antioquia, in the northern part of the Colombian Central Cordillera. Daedalma drusilla tinta occurs sympatrically with a local population of D. inconspicua , associated provisionally with the widespread subspecies orientalis n. ssp. Daedalma drusilla tinta is most commonly found slightly below the top of the San Felix peak at 2800–3000 m, whereas D. inconspicua was collected on the summit. Daedalma inconspicua is found consistently above 3000 m throughout its range, whereas all known individuals of D. drusilla were collected below 3000 m, suggesting a parapatric elevational replacement pattern for these two species.