Daedalma dinias dinias Hewitson

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 9-10

Daedalma dinias dinias Hewitson

( Figs. 1A, 1B, 8A, 13A)

Daedalma dinias Hewitson ; Butler, 1868: 183; Kirby, 1871: 107; Maassen & Weymer, 1890: 33; Thieme, 1906: 134; Weymer, 1912: 266, pl. 56, row f (venter), dorsum (misidentification of D. boliviana ); Gaede, 1931: 509; Adams, 1986: 252–253.

Daedalma dinias var. oenotria Weymer, 1912: 266 , pl. 56, row f (venter). Syntype series, Colombia: Bogotá [not located]; Gaede, 1931: 509. n. syn.

Material examined: COLOMBIA: LECTOTYPE female: Bogotá (Child), Rothschild Bequest, B.M. 1939-1, Type No. Rh. 4063, BMNH (designated by Adams, 1986: 252); 1 female: Bogotá (Child), Rothschild Bequest, B.M. 1939-1, BMNH; 1 female: Felder Coll., Rothschild Bequest, B.M. 1939-1, BMNH; 1 female: Bogotá, Rothschild Bequest B.M. 1939-1, BMNH; 1 female: Columbien, ex coll. H. Fruhstorfer, Fruhstorfer coll. 1933-31, BMNH; 1 female: Col., 150, Joicey Bequest B. M. 1934-120, BMNH; 1 female: Interior of Colombia, J. Carder, Joicey Bequest B. M. 1934-120, BMNH; 1 male: S. Martín, Llanos of Río Meta, G. D. Child, Godman-Salvin Coll., 1904.-1, BMNH; 1 male: Pacho , Chapman , Godman-Salvin Coll., 1904.-1, BMNH; 1 male: Nouv. Grénade, de Bogotá a Buenaventura, Dr. O. Thieme, 14.Dec.[18]77 a 22.Fev.[18]78, ex Oberthür Coll., Brit. Mus. 1927-3, BMNH; 1 male: Bogotá, 89-154, BMNH; 2 males: Boyacá, Arcabuco , 2300 m, 01.VII.1990, J-F. Le Crom leg., JLCB; 4 males: Boyacá, Arcabuco, 12.VIII.1989, J-F. Le Crom leg., JFLB; 1 male: Columbia, collection W. Schaus, USNM; 3 males: Bogotá, E. A. Smyth collection 1947, USNM; 1 male: Pacho, Cundinamarca, ex coll. Staudinger & Bang-Haas, TWP; 1 male: Chanchamayo, Peru [mislabeled], Rothschild Bequest 1939-1, BMNH.

Redescription: MALE ( Fig.1A): FWD dark brown; an oval reddish orange postmedian patch extending from mid-discal cell to cell Cu1-Cu2. HWD uniform dark brown. FWV blackish brown; subapical, apical and marginal area to vein Cu1 chestnut, chocolate brown and dusty yellow with two blackish rounded dots in R5-M1 and M1- M2; brick-red oval patch of FWD not visible. HWV with a nondescript pattern of chocolate brown, chestnut and dusty white patches and lines, with two prominent silver submarginal patches, one roughly triangular in M2-M3 contiguous to vein M3, and another, lunular in M3-Cu1. Genitalia ( Fig. 8A): Uncus short and stout; gnathos 2/3 length of uncus, stout; valvae oblong, with a conical distal dorsal process pointing upwards; saccus very long but slightly shorter than in leticia n. ssp.; aedeagus length of valva + saccus, very slightly contorted towards apical end, smooth.

FEMALE ( Fig. 1B): FWD ground colour medium brown, lighter distally; oblique orange band, lighter and wider than in male, extending to costa; three small subapical whitish patches dusted with grey; apical area suffused with grey. HWD medium brown, slightly lighter than in male with diffused orange subapical patches. FWV ground colour slightly lighter brown than in male; oblique band showing from upperside, diffused; subapical whitish scaling not apparent in male. HWV similar to male with more prominent yellow scaling. Genitalia ( Fig. 13A): Papillae anales flattened and hairy, their outer wall homogeneous without a bump; apophyses posteriores rudimentary, marked as short triangular sclerotized plates. Sinus vaginalis very deep, conical in shape in ventral view; antrum very short, connecting with sinus vaginalis through cleft-like opening; ventral wall of sinus vaginalis-lamella antevaginalis with parallel wrinkles; lamella postvaginalis with broadly bent, V-shaped batten pointing ventrally and stretching above sinus vaginalis; bursa copulatrix rounded in ventral view, with a pair of ribbon-like signa consisting of minute cuticular teeth; ductus bursae shorter than length of bursa, asymmetrically connecting with corpus bursae.

Remarks: Pyrcz (2004) pointed out that boliviana Staudinger , considered by many ( Weymer, 1912; Forster, 1964; Adams, 1986 and Lamas et al., 2004) as the southern subspecies of Daedalma dinias , actually represents a separate species. In fact, the markings of the HWV and male genital morphology of D. dinias suggest a closer affinity with the west Andean D. parvomaculata and D. eliza n. sp. than with D. boliviana . The original figures in Hewitson’s description represent a male (dorsal) and a female (dorsal and ventral) belonging to two species. Adams (1986) designated as the lectotype of D. dinias a female syntype bearing the label “ Bogotá ”. He justified his choice by indicating that Hewitson described dinias from Colombia, and that the female syntype comes from Colombia, whereas the other syntype, a male, is from Bolivia or Peru. This proves correct. Adams, however, went on to imply that the type female is from the Colombian Central Cordillera, rather than the Eastern Cordillera (Bogotá area), which is wrong. Adams held that the Bogotá population, described as D. dinias form oenotria Weymer (1912), is distinguished by an all brown FWV. He failed to notice that this is true only for males, whereas in females the orange oblique patch does show on the ventral surface. A comparison of several females, from both the Central and Eastern Cordilleras, demonstrates that the female lectotype is from the Bogotá region, as correctly indicated on the label. Consequently, oenotria is here considered a junior synonym of nominate D. dinias . Misled by the original error in the Hewitson plates, Weymer (1912) produced another synonym. He synonymized boliviana with the nominate dinias , and described the form rhomboidea of D. dinias from Bolivia (original illustration of ventral surface). This taxon was synonymized with nominate D. boliviana by Pyrcz (2004). Daedalma dinias dinias is found exclusively in the Colombian Eastern Cordillera, on both the western (Boyacá, Cundinamarca) and eastern slopes of its range (Meta, Caquetá). It flies in mid-elevation cloud forest at 2200–2600 m. It is uncommon and occurs in small, localized populations.