Daedalma inconspicua orientalis Pyrcz, 2011

Pyrcz, Tomasz W., Greeney, Harold F., Willmott, Keith R. & Wojtusiak, Janusz, 2011, 2898, Zootaxa 2898, pp. 1-68 : 34-35

Daedalma inconspicua orientalis Pyrcz , n. ssp.

( Figs. 2G, 5G, 5H, 12A, 17A, 21C, 21D, 21F, 21H, 22A, 23D)

Material examined: ECUADOR: HOLOTYPE male: Tungurahua, Tungurahua volcano, above Pondoa , 3200– 3250 m, I.2002, J. Wojtusiak, R. Garlacz and I. Aldaz leg., red, rectangular label saying : Holotype, MZUJ; PARA- TYPES (53 males and 6 females) : 6 males: Tungurahua, Tungurahua volcano, above Runtún , 2600–3000 m, 06.VIII.1998, T. Pyrcz leg., TWP (4 males to be deposited in MECN) 1 male: same locality, 3000 m, VIII.1998, I. Aldaz leg., TWP ; 1 male: Tungurahua, Baños , V.1998, I. Aldaz leg., TWP ; 1 male: Tungurahua, Tungurahua volcano, above Pondoa , 3300 m, 12.I.2002, J. Wojtusiak leg., TWP ; 1 male: same data but 3200–3250 m, I. Aldaz leg., TWP ; 5 males: Tungurahua, Morogacho , 3400–3500 m, IX.2004, O. Velástegui leg., TWP ; 1 male: Tungurahua, El Tablón, 3050–3100 m, 17.II.2004, T. Pyrcz leg., TWP ; 1 male: same locality, I.2005, O. Velástegui leg., TWP ; 2 males: same locality, I.1999, I. Aldaz leg., TWP ; 1 male: same locality, II.1999, I. Aldaz leg., TWP ; 1 male: same locality, 2600–3000 m, TWP ; 1 male: same locality, VIII.1998, I. Aldaz leg., TWP ; 1 male: Tungurahua, Triunfo-Patate , 26 km de Baños, 3000 m, 21.XI.1998, P. Boyer leg., TWP ; 2 males: Tungurahua, El Tablón, Triunfo-Patate , 26 km de Baños, 3000 m, VIII.1998, E. Aldaz leg., PBF ; 1 male: Tungurahua, El Tablón, Baños vers Patate km 18, 3000 m, 13.II.2004, P. Boyer leg., PBF ; 1 male: Tungurahua, La Cima, km 28 Ulba-Patate rd. , 3200 m, xii.1997, I. Aldaz leg., KWJH ; 2 males: Tungurahua, Runtún , 3000 m, VIII.1998, E. Aldaz leg., PBF ; 1 female: Tungurahua, Volcán Tungurahua, above Pondoa , 3200–3250 m, I.2002, I. Aldaz leg., TWP ; 1 female: same locality, 19.I.2001, J. Wojtusiak leg., TWP ; 1 female: Tungurahua, Tungurahua volcano, above Runtún , 3100 m, 06.VIII.1998, T. Pyrcz leg., TWP ; 1 female: Tungurahua, El Tablón, 3050–3100 m, 17.II.2004, T. Pyrcz leg., TWP ; 10 ♂: Tungurahua, Baños, Runtún , 3000 m, VIII.1998, E. & I. Aldaz leg., MBLI ; 5 males: same data but III.1999, MBLI ; 2 males: Tungurahua, Baños , El Tablón, 3000 m, VIII.1988, I. Aldaz leg., MBLI ; 5 males: Tungurahua, Pondoa , 3200–3400 m, I.2002, I. Aldaz leg., MBLI ; 1 female: Tungurahua, Tungurahua volcano, above Pondoa , 3200–3250 m, I.2002, I. Aldaz leg., TWP . Additional material: 1 male: Sucumbíos, Julio Andrade, Huaca-La Bonita km 45, 2500–2700 m, 09.V.2000, P. Boyer leg., PBF ; 1 female: Morona-Santiago, Chigüinda-Sigsig road, 3000 m, 03.XII.1998, P. Boyer leg., PBF ; 2 males: Morona-Santiago, km 17 Chigüinda-Gualaceo rd. , 2670 m, 14.X.2007, K. Willmott leg., FLMNH ; 1 female: Morona-Santiago, km 51.5 Cebadas-Macas rd., Quebrada Galgalán , 23.X.2007, K. Willmott leg., FLMNH . COLOMBIA: 1 male: Antioquia, San Felix , II.2002, 3050 m, G. Rodríguez leg., GRM ; 1 male: Colombia, Tolima, La Línea, 1987, G. Rodríguez leg., GRM ; 1 male: Tolima, La Línea, 3100 m, 03.V.1974, S. & L. Steinhauser leg., FLMNH .

Description: MALE ( Fig. 5G): Head, thorax and abdomen: not differing from nominate subspecies. Wings: FW (length: 26–28 mm, mean: 27.4 mm, n=30); differs from nominate subspecies by darker FW and HWD, and slightly larger size; FWV without any reddish suffusion contrary to nominate; lighter postdiscal area in M3-Cu1 and Cu1-Cu2 widely connected to lighter area, of similar colour, covering distal half of discal cell; HWV ground colour dark cholocate brown, lighter than in nominate and other west Andean subspecies, similar to tapichalaca n. ssp. Genitalia ( Fig. 12A): Uncus stout and nearly straight; gnathos half length of uncus, very stout basally, gradually narrowing to a sharply ended tip; saccus shorter than in other subspecies, except for onorei, similar in this respect; valvae shorter than in most other subspecies, similar to cuencana, with a smooth ampulla; aedeagus not differing from other subspecies.

FEMALE ( Fig. 5H): FW length: 28–32 mm, mean: 29.8 mm, n=5; FWD without any reddish or orange markings, contrary to variegata or nominate; HWD with reddish suffusion covering postdiscal to submarginal area, somewhat variable. Genitalia ( Fig. 17A): Sinus vaginalis widely rounded. Long axes of ductus bursae and antrum running under wide angle in relation to each other, so that corpus bursae points anterio-ventrally. Antrum short. Colliculum well pronounced, with two ring-like sclerotizations. Ventral wall of ninth segment posteriorly to papillae anales with strongly marked, parallel wrinkles. Sclerotized lateral parts of eighth segment that connects tergum with sinus vaginalis wide. Posterior surface of V-shaped batten on lamella postvaginalis covering entrance to anterior parts of sinus vaginalis, wide, with triple bends. Bursa copulatrix with two ribbon-like signa, consisting of minute teeth. This subspecies differs from D. inconspicua palacio and D. inconspicua variegata by sharply bent central part of hump on lamella postvaginalis.

Etymology: The name orientalis (eastern) refers to the distribution of this subspecies in the eastern Andes of Ecuador.

Remarks: This subspecies has the widest distribution of all D. inconspicua taxa, being known in Colombia in both the Cordilleras Occidental (Nariño and Caldas) (Salazar, 2004) and Central as far north as Antioquia, and in eastern Ecuador from Sucumbíos to Morona-Santiago. The few known Colombian individuals do not appear to differ from east Ecuadorian orientalis, but nevertheless were not included in the type series, as further collecting may prove that they do in fact represent distinct subspecies. An individual from Tolima is shown in Fig. 2G.