Copris (Paracopris), Balthasar, 1939

Bui, Van Bac, Ziegler, Thomas & Bonkowski, Michael, 2019, Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 4712 (1), pp. 51-64 : 54

Subgenus Paracopris Balthasar, 1939

Type species Copris punctulatus Wiedemann, 1823 (by original designation)

Nomenclatural history. Recently, the subgenus Paracopris was considered as a genus by Sabu et al. (2011), Král & Bezděk (2016) and Cheung et al. (2018), but a lack of the genetic analysis makes the generic placement of Paraco- pris unclarified. In this study, we regarded Paracopris as a subgenus of the genus Copris according to the traditional classification of Balthasar (1939) that has been widely used in taxonomic studies.

Diagnosis. Color: grayish-black, reddish brown or chocolate-brown. Whole surface slightly convex to fairly flat. Head broad. Clypeal surface smooth (i.e. minutely punctate). Clypeus and frons typically armed with a horn and/or carina. Pronotum broad, anteriorly untruncated and distinctly punctate. Elytra distinctly striate; elytral intervals strongly punctate. Tibia, compared with the other subgenera slender. Metatibia bear a weak transverse outer carina. Metatarsus slender, and with five tarsomeres; metatarsomeres approximately equal in length.