Lomechusoides amurensis ( Wasmann, 1897 )

Jászay, Tomáš & Hlaváč, Peter, 2013, A taxonomic revision of the myrmecophilous genus Lomechusoides Tottenham, 1939 (Coleoptera: Staphylinidae: Aleocharinae) Part I. Redescription of the genus, definition of species groups and th, Zootaxa 3683 (1), pp. 65-81: 71-75

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Lomechusoides amurensis ( Wasmann, 1897 )


Lomechusoides amurensis ( Wasmann, 1897) 

( Figs. 14 –27View FIGURES 14 – 20. LView FIGURES 21 – 27. L, 49View FIGURES 48 – 50. 48, L)

Lomechusa amurensis Wasmann, 1897: 246 

Lomechusa Ganglbaueri Bernhauer, 1936: 323  . Type data: Ostsibirien: Irkutsk (im British Museum und in meiner eigenen Sammlung), Mongolei: Shangai (in der Sammlung des naturhistorischen Staatsmuseum in Wien), part, synonymy in Hlaváč et al., 2011: 14.

Lomechusa ganglbaueri  ssp. uralensis Bernhauer, 1936: 323. Type data: ein weiteres Exemplar vom Ural (Slatoust) [=Zlatousť], synonymy in Smetana, 2004: 457, synonymy confirmed.

Type material studied: Lomechusoides amurensis Wasmann  : HOLOTYPE, 13: (h) yellow parabolic label: Amur land / (h) Lomechusa  n. sp. nec inflata Thoms. ?? L. sibirica Motsch.  / (h) Lomechusa amurensis Wasm.  n. sp. Type / round label with red margin: (p) Type / red label: (p) Holotypus, (h) Lomechusa amurensis Wasmann, W. Schillow  des. 981 / (p) Sharp Coll 1905 - 323 / red label (p) HOLOTYPE 3 Lomechusoides amurensis (Wasmann)  , P. Hlaváč det. 2007. BMNH.

Lomechusoides ganglbaueri uralensis Bernhauer  : HOLOTYPE, 13: (h) Ural, Slatoust [=Zlatousť]/ (h) ganglbaueri Brh.  (p) det. Bernh. / orange label (h) ssp. uralensis Bernh. Typus unic. / (p) Chicago NHMus, M. Bernhauer Collection / red label (p) Lomechusa ganglbaueri  ssp. uralensis (Bernhauer), P. Hlaváč des., 2009 / Lomechusoides amurensis (Wasmann)  = Lomechusa ganglbaueri  ssp. uralensis (Bernhauer) P. H l av á č det., 2009. FMNHAbout FMNH.

Other material. 1 Ƥ: in Russian characters: Vost. [=eastern] Sayan 1500m, verkh [=top] Kyngarga, 18– 23.07. 95 / iz mkhov [from moss], Shavrin, A. / Lomechusa amurensis Wasm. Shavrin  der. 97, in toto in euparal.

CPHK. 1 3: in Russian characters: promysel Ozerpach liman Amura, 16 – 12.vi. 1915, Chernavin / L. amurensis W. Schilov  det. 19. ZINAbout ZIN. 1 Ƥ: in Russian characters: Syutszukte, yu-b. [south-east], Kenteys. Z, Urgi, Kozlov, 1.vi. 925. ZINAbout ZIN.

Redescription. Body lighter reddish-brown ( Fig. 49View FIGURES 48 – 50. 48, L); head and narrow basal margin of tergites darker, slightly shining.

Head ( Fig. 15View FIGURES 14 – 20. L) quadrate, longer than wide (HW/HL= 0.79), with deep median, frontal, triangular depression which is sparsely microsculptured, anterior part shining; length of eyes 0.26 times of length of head; temples slightly convergent posteriorly, posterior corners of temples rounded, posterior margin of head rounded, postoccipital proccess well-defined, protuberant. Antennae ( Fig. 16View FIGURES 14 – 20. L) shorter, scape 1.35 times as long as wide and 1.95 times as long as pedicel, all antennomeres elongate except II which is as long as wide, III 1.10 and IV 1.27 as long as wide; antennomeres IV–X elongate, oval, terminal segment thin and acute, 1.15 times as long as scape, relative length of antennomeres from base to apex: 21: 10: 13: 14: 17: 16: 16: 17: 16: 16: 24.

Pronotum ( Fig. 17View FIGURES 14 – 20. L) unicoloured, with clearly convex anterior margin, sides rounded with basal corners obtuse and with deep lateral depressions, surface of this depressions with microsculpture, disc smooth, with sparse, uneven puncturation, basal median lobe large; pronotum in hind corners as wide as humeral width of elytra; ratio: PW/PL = 1.42. Metaventrite ( Fig. 18View FIGURES 14 – 20. L a) with metaventral process ( Fig. 18View FIGURES 14 – 20. L b) with anterior corners sharply angled, anterior margin straight, lateral margins convex, metaventral process in anterior part with lens-like depression bounded with rounded groove, surface unevenly punctured and with short setae, setae not reaching margin of next puncture, metaventral disc lacking microsculpture, unevenly punctured, in places lacking punctures.

Elytra finely punctured, distance between punctures greater than puncture diameter, 1.15 times shorter than pronotum measured on median line, ratio: EW/EL=2.00.

Abdomen with tergite II densely punctured and setaceous, distance between punctures on posterior margin greater than puncture diameter or even double of diameter of puncture, tergites III–IV with fine puncturation and setation on posterior margin, tergites V–VII sparsely but finely punctured at base, densely punctured on lateral parts, tergites II–VIII with visible microsculpture on lateral sides, tergite and sternite VIII as in Figs. 19View FIGURES 14 – 20. L a, 19 b, 20 a, 20 b.

Aedeagus as in Figs. 21, 23, 24View FIGURES 21 – 27. L, median lobe robust, basal bulb wide with well-developed distal crest and basal bridge. Internal structures weakly-defined, apical lobe tube-like, apical part sharp, with many pseudopores. Paramere ( Fig. 22View FIGURES 21 – 27. L) with triangular paramerite, velum about as large as paramerite, apical lobe subparallel, hookshaped bent backwards, apex with about four setae. Spermatheca as in Figs. 25–27View FIGURES 21 – 27. L, basal part V-shaped, apical part as long as basal part, sligthly broader and 2.5 –3.0 times as long as wide, internal wall in apical part roughly wrinkled.

Measurements. (Holotype), body length 4.85 mm; forebody length 2.40 mm; head length 0.95 mm; head width 0.75 mm; antennal length 2.60 mm; pronotal length 1.10 mm; pronotal width 1.57 mm; elytral length on sutura 0.85 mm; elytral width 1.70 mm; foretibial length 1.00 mm; midtibial length 1.20 mm; hindtibial length 1.50 mm.

Differential diagnosis. L. amurensis  can be distinguished from other species of the group by the characteristic shape of the metaventral process with deep median depression and by the shape of spermatheca.

Host ant. Unknown

Distribution. Russia (Ural, Siberia and Far East) and Mongolia

Lomechusoides schneideri Maruyama et Hlavá  č, 2004 ( Figs. 28 –31View FIGURES 28 – 31. L, 50View FIGURES 48 – 50. 48, L)

Lomechusoides schneideri Maruyama & Hlaváč, 2004: 110  . Type data: China m., 19 –VII– 1992, Sichuan, 3,600 m, Kangding env.

Type material studied. HOLOTYPE, 1 Ƥ: (p) China m., 19.7. 1992, Sichuan, 3600m, Ken ding env., J. Schneider legit. / (p) ex coll. M. Dvořák, National Museum Prague, Czech Republik/ red label (p) Holotype, Lomechusoides schneideri  des. Maruyama & Hlaváč, 2004.

Supplementary description. The species was recently adequately described by Maruyama & Hlaváč (2004: 110). Head ( Fig. 28View FIGURES 28 – 31. L), pedicel as long as wide, antennomere III about 1.4 times as long as wide, both equal in length, relative length of antennomeres from base to apex: 22: 11: 12: 14: 15: 16: 15: 15: 15: 14: 20. Pronotum ( Fig. 29View FIGURES 28 – 31. L) with sides and medium line with dense microsculpture, dull, pronotal disc near medium line with fine microsculpture, glossy.

Pronotum in hind corners wider than humeral width of elytra. Metaventrite with metaventral process with anterior corners sharply angled, anterior margin slightly convex, lateral margins convex ( Fig. 30View FIGURES 28 – 31. L), metaventral process in anterior part lacking shalow depression and rounded groove, whole surface of metaventrite unevenly but densely punctured and with short setae, setae reaching margin of next puncture, metaventral disc with sparse microsculpture. Basal part of elytra densely punctured, distance betwen puntures shorter than diameter of puncture, densely and markedly microsculptured, dull, rest of surface of elytra coarsely punctured, lacking microsculpture, glosy. Abdomen with tergite II densely punctured and setaceous, distance between punctures on posterior margin as long or slightly longer than puncture diameter, posterior margin of tergites III–IV with fine, sparse puncturation and setation, tergite IV much sparsely punctured than III, tergites V–VI sparsely, finely and evenly punctured, tergites II–VI in basal part with fine microsculpture, tergite and sternite VIII as in Figs 31View FIGURES 28 – 31. L a, 31 b.

Diferencial diagnosis. L. schneideri  can be easily distinguished from the other species of the L. amurensis  group by the long and quadrate head and long, subparallel temples. Host ant. Myrmica cf. ruginodis Nylander, 1846 

Distribution. China (Sichuan)


Field Museum of Natural History


Russian Academy of Sciences, Zoological Institute, Zoological Museum














Lomechusoides amurensis ( Wasmann, 1897 )

Jászay, Tomáš & Hlaváč, Peter 2013


Lomechusa Ganglbaueri Bernhauer, 1936 : 323

Hlavac 2011: 14Bernhauer 1936: 323


Lomechusa ganglbaueri

Smetana 2004: 457
Bernhauer 1936: 323


Lomechusa amurensis

Wasmann 1897: 246


Lomechusoides schneideri Maruyama & Hlaváč, 2004 : 110

Maruyama 2004: 110