Pronephrium stiriacum (UNGER) E. KNOBLOCH et KVAČEK

Kvaček, Zlatko & Teodoridis, Vasilis, 2011, The Late Eocene Flora Of Kučlín Near Bílina In North Bohemia Revisited, Acta Musei Nationalis Pragae Series B 67 (3 - 4), pp. 83-144 : 87

Pronephrium stiriacum (UNGER) E. KNOBLOCH et KVAČEK

Pl. 1, figs 6, 10

1847 Polypodites styriacus UNGER ; p. 121, pl. 36, figs 1-5.

1866 Aspidium fischeri auct. (non HEER); Ettingshausen, p. 17, pl. 3, figs 9-11.

1852 Goniopteris stiriaca (UNGER) A. BRAUN , p. 553.

1855 Lastraea stiriaca (UNGER) HEER , p. 31, pls 7-8.

1960 Cyclosorus stiriacus (UNGER) CHING et TAKHTAJAN in Fataliev, p. 1217.

1962 Cyclosorus stiriacus (UNGER) GRAMBAST , nom. illegit, p. 24, text-fig. 4.

1963 Abacopteris stiriaca (UNGER) CHING , p. 298.

1976 Pronephrium stiriacum (UNGER) E. KNOBLOCH et KVAČEK , p. 12, pl. 1, fig. 1.

2002a Pronephrium stiriacum (UNGER) E. KNOBLOCH et KVAČEK ; Kvaček, p. 221, pl. 4, fig. 7.

Incomplete sterile pinnae more than 50 mm long and 14 mm wide with a stout midrib, crenate on sides according to almost fused pinnules, incisions between pinnules 2–3 mm, pinnules max. 7 mm long, at angles of 55–60°, at the apex rounded and slightly turned upwards, midrib of pinnules almost the same thickness as secondaries, straight to slightly bent on very end, secondaries in 6 pairs on each side, the basal united between adjacent pinnules (goniopterid), high- er slightly bent, merging the margin.

D i s c u s s i o n: Leaf fragments of this common Tertiary swamp fern, showing a goniopterid venation (pl. 4, fig. 7), were wrongly assigned to Aspidium due to compression of the specimens and poor preservation at Kučlín by Ettingshausen (1866) and assigned to a different morphotype. In one of the specimens (BP. 55.2319.1, not illustrated by Ettingshausen) and a new material, the goniopterid venation is clearly seen. The specimens from Kučlín are very rare and indeed more delicate than the type material from the Miocene of Styria (Schöneck, Unger 1847, pl. 36, figs 1-4) and other places in the European Tertiary (e.g., Barthel 1976), probably due to different non-swampy environments. Pronephrium penangianum (HOOK.) HOLTTUM (syn. Abacopteris penangiana (HOOK.) CHING ) from the Himalayas is considered according to several authors (e.g., Ching 1963) as the nearest living relative.

M a t e r i a l s t u d i e d: BP 55.2319.1, BP 55.2353.1

(two specimens on the same slab), DB KUC 234 .

? Lomariopsidaceae