Zodarion maculatum ( Simon, 1870 )

Benhalima, Souâd & Bosmans, Robert, 2020, Revision of the genus Zodarion Walckenaer, 1833 (part IV). The species of Morocco (Araneae: Zodariidae), Zootaxa 4899 (1), pp. 93-114 : 99-102

Zodarion maculatum ( Simon, 1870)

Figures 25–26 , 34–38 , 49–53

Enyo maculata Simon, 1870: 146 (Lectotype ♀ and paralectotype ♀, designated by Bosmans 1994, deposited in MNHN:

ITALY: Sicily—examined).

Zodarium maculatum: Denis 1935: 67 , fig. 7; Denis 1937: 14.

Zodarion variegatum Denis, 1956: 201 , figs 12–14 (Holotype ♀, deposited in MNHN: MOROCCO: Casablanca, Aïn Sbaâ, leg. Gattefossé, 2.VIII.1951 —examined). syn. nov.

Zodarion maculatum: Bosmans 1994: 127 , figs 39–41, 103–104.

Remarks. The tube with the type of Zodarion variegatum includes a label Zodarion fusciventris , with Denis’ handwriting, but a species with this name has never been described. Moreover, the specimen fits completely Denis’ (1956) description and the locality is the same as the type locality of Z. variegatum , so it is obviously the holotype of Z. variegatum .

Examination of the holotype female of Z. variegatum and comparison with our material of Z. maculatum shows they are the same species. Denis’ excellent fig. 13 of the epigyne of Z. variegatum shows clearly the postero-median process in the epigynal pit, as mentioned in the diagnosis below.

Diagnosis. The species cannot be mistaken for any other species of the elegans group by the long, curved and terminally pointed tibial apophysis in the male ( Fig. 34 ), and by the shape of the rounded epigynal pit with a posteromedian triangular process in the female ( Fig. 50 ).

Description. See Bosmans (1994), and Figs 25–26 , 34–38 , 49– 53 in the present paper.

Previous records. MOROCCO: Casablanca-Settat, Casablanca, Ain-Sbaâ, 33°37’45’’N, 07°29’56’’W, leg. Gattefossé, 2.VIII.1951, 6Ƌ, 1♀ ( Denis 1956; MNHN, holotype of Z. variegatum ); Marrakech-Safi, Marrakech, Oued Tensift, 31°41’33’’N, 07°59’17’’W, IV.1930, 1♀ (Caporiaco 1932); Tanger-Tétouan-Al Houceima, Tanger, 35°37’00’’N, 05°47’00’’W, E. leg. Simon, 1868, 2♀♀ ( Simon 1870; MNHN).

Other material examined. None.

Distribution. NW Morocco, Algeria, S Spain, S Italy and Sicily (Map 2).

Figures

FIGURES 23–28. Dorsal habitus of Zodarion valentii Bosmans, Loverre & Addante, 2019 male (23) and female (24), Z. maculatum (Simon, 1870) male (25) and female (26), and Z. maghrebense sp. nov. male (27) and female (28).

FIGURES 29–43. Genitalic morphology of Zodarion valentii Bosmans, Loverre & Addante, 2019 (29–33), Z. maculatum (Simon, 1870) (34–38) and Z. maghrebense sp. nov. (39–43). 29, 30, 34, 35, 39, 40 Male palp, ventral view; 31 Tibial apophysis, retrolateral view; 32, 33, 37, 38, 42, 43 Male palp, retrolateral view; 36, 41 Tip of embolus, ventral view.

FIGURES 44–58. Genitalic morphology of Zodarion valentii Bosmans, Loverre & Addante, 2019 (44–48), Z. maculatum (Simon, 1870) (49–53) and Z. maghrebense sp. nov. (54–58). 44, 45, 49, 50, 54, 55 Epigyne, ventral view; 46, 48, 51, 53, 56, 58 Vulva, ventral view; 47, 52, 57 Vulva, dorsal view.