Polynoncus burmeisteri Pittino, 1987

Costa-Silva, Vinícius, Strümpher, Werner P., Thyssen, Patricia J. & Vaz-de-Mello, Fernando Z., 2024, Taxonomic revision of the South American genus Polynoncus Burmeister, 1876 (Coleoptera: Scarabaeoidea: Trogidae), Journal of Natural History 58 (1 - 4), pp. 14-166 : 61-62

Polynoncus burmeisteri Pittino, 1987

( Figures 19–20 )

Polynoncus burmeisteri Pittino, 1987: 380 (original description); Scholtz 1990: 1431 (redescription); Gómez 2008: 516 (key to Argentinean species); Zidek 2013: 8 (catalogue); Zidek 2017: 98 (catalogue); Costa-Silva and Diéguez 2020: 271 (records from Argentina)

Type specimens examined. HOLOTYPE. (♂ MNHN – Figure 20 ) First label [white, aged, Edgar von Harold’s handwriting): ‘Mendoza’. Second label [light brown with black border, typeset]: ‘EX MUSEO / BONVOULOIR’. Third label [white, aged, with black border, typeset]: ‘MUSÉUM PARIS / 1936 / COLL. A. BOUCOMONT’. Fourth label [red, Riccardo Pittino’s handwriting]: ‘ Polynoncus / burmeisteri n. sp. / HOLOTYPUS / Det. R. Pittino 1987 ’. Fifth label [white with black border, typeset]: ‘MNHN / EC10487’ ( Figure 20e ). Type locality: ‘ Mendoza’, Argentina.

PARATYPES (1 ♂ and 3 ♀ RPMI [through detailed photographs]; 2 ♂ and 2 ♀ MNHN) .

First label [white, typeset]: ‘Bella Vista / Arg., Gassner’. Second label [white, typeset]: ‘EX COLL. / WINKLER, WIEN’. Third label [red, typeset]: ‘PARATYPUS’. Fourth label [red with black border, Riccardo Pittino’s handwriting]: ‘ Polynoncus ♀ / burmeisteri / n. sp. / Det. Pittino 1987 ’ (1 ♀ RPMI). First label [green, handwritten]: ‘Bolivia’. Second label [white, typeset]: ‘MUS. LA FERTÉ / 380’. [handwritten]. Third label [light brown with black border, typeset]: ‘Ex-Musӕo / D. Sharp 1890’. Fourth label [green with black border, typeset]: ‘Muséum Paris / ex Coll. / R. Oberthür / 1952’. Fifth label [red, typeset]: ‘PARATYPUS’. Sixth label [red with black border]: ‘ Polynoncus / burmeisteri n. sp. / Det. Pittino 1987 ’ (1 ♀ RPMI). First label [white, aged, handwritten]: ‘Mendoza’. Second label [green, typeset]: ‘Museum Paris / ex Coll. / R. Oberthur’. Third label [red, typeset]: ‘PARATYPUS’. Fourth label [red with black border, Riccardo Pittino’s handwriting]: ‘ Polynoncus ♀ / burmeisteri n. sp. / Det. Pittino 1987 ’ (1 ♀ RPMI). First label [white, aged, handwritten]: ‘Mendoza’. Second label [white]: ‘EX COLL. / WINKLER, WIEN’. Third label [red with black border, typeset]: ‘PARATYPUS’. Fourth label [red with black border, Riccardo Pittino’s handwriting]: ‘ Polynoncus / burmeisteri ♂ / n. sp. / Det. Pittino 1987 ’ (1 ♂ RPMI). First label [white, aged, handwritten]: ‘Mendoza’. Second label [light brown with black border, typeset]: ‘Ex Musaeo / E. Steinheil’. Third label [white, handwritten]: ‘ ♀ ’. Fourth label [light green with black border, typeset]: ‘Muséum Paris / ex Coll. / R. Oberthür / 1952’. Fifth label [red, Riccardo Pittino’s handwriting]: Polynoncus ♀ / burmeisteri n. sp. / Det. R. Pittino 1987 ”. Sixth label [red, Riccardo Pittino’s handwriting]: ‘PARATYPUS’. Seventh label [white with black border, typeset]: ‘MNHN / EC10488’ (1 ♀ MNHN). First label [white, aged, handwritten]: ‘gemmifer / mendoza’. Second label [green with black border, typeset]: ‘Museum Paris / ex Coll. / R. Oberthur’. Third label [red with black border, typeset]: ‘PARATYPUS’. Fourth label [red with black border]: ‘ Polynoncus ♀ / burmeisteri / n. sp. [Riccardo Pittino’s handwriting] / Det. Pittino 1987 ’. Fifth label [white with black border]: ‘MNHN / EC10489’ (1 ♀ MNHN). First label [ligh brown, handwritten]: ‘gémifer / Blanchard. / Mendoza’. Second label [white, aged, with black border, typeset]: ‘Ex-Musae / A. SALLÉ / 1897’ [vertical, right side]. Third label [light green with black border, typeset]: ‘Muséum Paris / ex Coll. / R. Oberthür / 1952’. Fourth label [white, aged, Patricia Vaurie’s handwriting]: ‘ Trox / gemmifer / Bl. / Vaurie ‘61’. Fifth label [red with black border]: ‘PARATYPUS’. Sixth label [red, Riccardo Pittino’s handwriting]: ‘ Polynoncus ♂ / burmeisteri n. sp. / Det. R. Pittino 1987 ’. Seventh label [white with black border]: ‘ Polynoncus / burmeisteri / Pittino ♂ [Andrew Smith’s handwriting] / Det: A.B.T. Smith 2016 [typeset]’. Eighth label [white, typeset]: ‘MNHN / EC10490’ (1 ♂ MNHN). First label [light brown with black border]: ‘MUSEUM PARIS / PATAGONIE / AIVES DU RIO NEGRO / H. DE LA VAULX 1897’. Second label [red with black border, typeset]: ‘PARATYPUS’. Third label [red with black border]: “ Polynoncus / burmeisteri ♂ / n. sp. [Riccardo Pittino’s handwriting] / Det. Pittino 1987. Fourth label [white with black border]: ‘MNHN / EC10491’ (1 ♂ MNHN).

Differential diagnosis. Polynoncus burmeisteri is similar to P. pseudogemmifer sp. n. and P. argentinus . However, the last two species have second elytral costae with well-developed tubercles, while P. burmeisteri has just a granulate row (see Costa-Silva and Diéguez 2020, p. 270, fig. 12).

Polynoncus burmeisteri also bears some resemblance to P. gemmifer , but the absence of setae in the hypomeron easily separates it from the latter (see section above). The male genitalia of the four species are very distinctive. For details of description and differential diagnosis, see Pittino (1987, p. 380) and Costa-Silva and Diéguez (2020, p. 269), respectively.

Geographic distribution. Polynoncus burmeisteri is recorded from central Argentina, with a single record from Bolivia (one paratype without specific locality) ( Figure 19 ).

Examined non-type material (14 specimens). ARGENTINA: Buenos Aires, Carmen de Patagones , no date or collector (1♂ MLPA) . Córdoba, Cabana, 22 December 1925, no collector (1♂ MLPA); Calamuchita, Hotel El Sauce , October 1975, M . Viana leg . (1♂ CVMD); Calamuchita, Hotel El Sauce , November 1968, M . Viana leg . (1♂ CVMD); Tanti , February 1943, M . Biraben leg . (1♂ MLPA) . Mendoza, Santa Rosa, Reserva de Biosfera de Ñacuñán , 11–13 December 2002, V .M . Diéguez leg . (1♂ and 1♀ CVMD); Santa Rosa, Reserva de Biosfera de Ñacuñán , 11–13 December 2002, V .M . Diéguez leg . (2♂ and 1♀ IADIZA); Santa Rosa, Reserva de Biosfera de Ñacuñán , 11–13 December 2002, V .M . Diéguez leg . (1♂ and 1♀ CEMT) . Río Negro, Aguada Cecílio, no date or collector (2♀ MLPA) .

Figures

Figure 19. Distribution map of Polynoncus bullatus (red circles) and P. burmeisteri (green triangles) in South America.

Figure 20. Holotype of Polynoncus burmeisteri Pittino, 1987: habitus in a, dorsal and b, lateral view; male genitalia in c, dorsal and d, lateral view; e, labels. Photos by Christophe Rivier (MNHN).