Martensodesmus cattienensis, Golovatch, Sergei I., Geoffroy, Jean-Jacques, Stoev, Pavel & Spiegel, Didier Vanden, 2013

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Stoev, Pavel & Spiegel, Didier Vanden, 2013, Review of the millipede family Opisotretidae (Diplopoda, Polydesmida), with descriptions of new species, ZooKeys 302, pp. 13-77 : 47-48

Martensodesmus cattienensis sp. n. Figs 27-29

Martensodesmus sp. - Golovatch et al. 2011b: 81.

Type material.

Holotype ♂ (MNHN JC 343), Vietnam, Dongnai Prov., Cat Tien National Park, lowland semi-deciduous tropical monsoon forest, ca 150 m a.s.l., 107°10'-107°34'E, 11°21'-11°48'N, 08-22.11.2005, leg. A. E. Anichkin.

Paratypes.

1 ♂, 1 ♀ (MNHN JC 343), 1 ♂, 1 ♀, 1 ♀ fragment (ZMUM), 1 ♂ (ZMUC), same locality, together with holotype; 1 ♂, 1 ♀, 1 ♀ subadult (MNHN JC 343), 1 ♂, 1 ♀ (SEM), 1 ♂, 1 ♀, 1 ♀ subadult (ZMUM), same locality, 01.06.2005, leg. A. E. Anichkin; 1 ♂ (MNHN JC 343), 1 ♂ (NMNHS) same locality, 15.07.2005, leg. A. E. Anichkin.

Diagnosis.

Differs readily fromcongeners by the missing modifications on the ♂ vertex, coupled with quite well developed shoulders on metaterga, the high and broad paraterga, as well as the presence near the gonopod telopodite’s midpoint of three strong spines proximally to a considerably attenuating acropodite.

Name.

To emphasize the type locality.

Description.

Length of holotype ca 5.5 mm, width of midbody pro- and metazona ca 0.8 and 1.0 mm, respectively. Length of ♂ paratypes ca 5.5-6.0 mm, width of midbody pro- and metazona ca 0.75-0.8 and 1.0-1.1 mm, respectively. Length of ♀ paratypes ca 7.0-8.0 mm, width of midbody pro- and metazona ca 0.9-0.95 and 1.1-1.15 mm, respectively. Coloration in alcohol from nearly uniformly light yellowish to head and several anterior segments slightly infuscate, light yellow-brown, more rarely with a rusty reddish tint.

Body with 19 (♂) or 20 (♀) segments. All characters like in Retrodesmus cavernicola sp. n., except as follows.

Antennae very long, but strongly clavate (Fig. 27G), extending behind segment 4 (♂) or 3 (♀) when stretched dorsally.

In width, collum << head = segments 2 & 3 <4 <5 <6=15 (16); thereafter body gradually tapering towards telson. Paraterga well-developed (Fig. 27), starting from a broadly rounded, kidney-shaped collum, mostly only very faintly declivous to continue the outline of a rather slightly convex dorsum, largely set high, at about ¼ of midbody height, with quite strong shoulders frontolaterally (Figs 27D, E). Caudal corner of postcollum paraterga mostly dentiform, always narrowly rounded and extending increasingly well behind rear tergal margin only in segments 15-18 (♂) or 16-19 (♀). Lateral edge of paraterga with 2 or 3 small setigerous indentations in poreless and poriferous segments, respectively. Ozopores very evident, round, flush open on dorsal surface, clearly removed from caudal margin and lying anteriorly to bottom of caudalmost lateral incision (Figs 27 A–C, E, F, H, I, L), lateral tooth being clearly shorter than medial one. Each postcollum metatergum with 3+3, long, bacilliform setae arranged in three regular transverse rows; polygonal bosses flat, but visible, while transverse sulcus mostly rather deep (Figs 27 A–I).

Legs rather long and slender, ca 1.4-1.5 (♂) or 1.2-1.3 (♀) times as long as midbody height (♂); tarsi longest and particularly slender (Figs 27A, B, G–I, K, 28D), sphaerotrichomes or other modified setae missing.

Gonopod telopodite (Figs 28 E–I, 29) clearly curved, but stout, unipartite; basal half voluminous and supplied with three strong spines, distal half gradually attenuating, apical piece (a) distal to a vestigial solenomere with a number of short spinules. Neither bacilliform ornamentations nor an accessory seminal chamber, nor a hairy pulvillus.

Remarks.

This species has already been referred to as Martensodesmus sp. elsewhere ( Golovatch et al. 2011b).