Homoneura pseudograndis, Papp & Gaimari, 2013

Papp, L. & Gaimari, S. D., 2013, The Holotype Of Homoneura Grandis (Kertész, 1915) With Description Of A New Species From Taiwan (Diptera, Lauxaniidae), Acta Zoologica Academiae Scientiarum Hungaricae 59 (1), pp. 31-39 : 35-39

Homoneura pseudograndis sp. n.

(Figs 4–6, 9–10, 12, 15)

Holotypemale (HNHM, onaNo. 1 collectionpin): 1) FormosaSauter; 2) Mt. Hoozan 1910. “V.”; [Kertész’shandwriting] “grandisKert.” Det. Kertész. Inagoodstateofpreser- vation; itshindlegsaswellasabdomenwithgenitaliaareprepared, andnowiskeptina microvialwithglycerolonthesamepin.

Measurementsinmm: bodylength 7.04, winglength 6.96, wingwidth 2.75. Figs 7–12. Homoneura species , malegenitalia. 7–8 = H. grandis (Kertész, 1915) , holotype: 7 = hypandrium, ventralview, 8 = medialpartofhypandriuminhighermagnification. 9–10 = H. pseudograndis sp. n., holotype: 9 = hypandrium (slightlyflattened), ventralview, 10 = medialpartofhypandriuminhighermagnification. 11–12 = phallus, lateralview: 11 = H. grandis (Kertész, 1915) , 12 = H. pseudograndis sp. n. Scales: 0.4 mm for Figs 7, 9, 11–12, 0.2

mm for Figs 8, 10.

Bodyasintheotherspeciesofthegroup. Longestaristalrays 0.20 mm. Subapical brown spots on R 2 +3, R 4 +5 and M 1+2 confluent, proximal spot on R 4 +5 0.75 mm long (in 4 of 5 specimens of H. grandis that is c. 0.50 mm) .

Malegenitalia. Cercus (Fig. 4) rounded, itslongestsetaeslightlylongerthan 0.2 mm. Hypandrium (Figs 9–10) stronglyasymmetricalwithanasymmetricaltriangularapodeme, n., holotype. Scales: 0.5 mm for Fig. 13 , 0.4 mm for Figs 14–15 .

lefthalfwithalongerprocess, whichbearsasmallsubapicaltooth, medialtoothjustonthe baseofleftprocess; righthalfwithadigitiformprocess, whichisslightlylongerthanin H. grandis . Surstyluslargerthanin H. grandis , caudalextensionbroad, with 2 smallanterior basalprojections; outer (lateral) surface (Fig. 5) withsimilarmarginalsetaeasin H. grandis , butothersetaefiner, inner (medial) surface (Fig. 6) withsmallsharppegsonitsapical 1/3 only. Postgonites ( Fig. 15 ) structurallythesameasin H. grandis (directconnectionofphallapodemetothemedialprocessonly, lateralprocessesconnectedtophallapodemethrough membranes). Innerprocessisthelongestone, straightandthin, armsofthelateralpairof processes ( Fig. 15 ) subequal, medialarmonlyslightlybroaderthanlateralone. Phallus (Fig. 12) withlesssharpenedbase, andslightlynarrowerapicalthirdthanin H. grandis . Phallapodemelongerandlesscurvedthanin H. grandis .

H. pseudograndis sp. n. isdifferentfrom H. grandis mainlyincharacteris-

ticsofmalegenitaliaasdescribedabove. Thisisthecasealsowithotherunde-

Figs 16–18. Homoneura grandis (Kertész, 1915) , holotype, male: 16 = adult, 17 = head, dorsal view, 18 = left wing. (Photos S. D. Gaimari)

scribedspeciesofthegroup. Itsspeciesdistinctnessisunderlinedbythefact thatitissympatricwith H. grandis (oralmostso; consideringthecomparatively smallareasizeofTaiwanbutknowingwelltheextremereliefofitsregions/ sceneries, onemustbecarefulinjudgingthiskindofquestion).

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Acknowledgements – WearegratefultoMs. LiShi (ShenyangUniversity, P. R. China) forinformationonherstudiesonthespeciesgroup, foradviceandforencouragingusto makethisstudy. OurthanksareduetoMrsTimeaPappandMrJánosPapp (Monor, Hungary) fortheirhelpinpreparingfigureplates.

Figures

Figs 13–15. Homoneura species, malegenitalia. 13 = H. grandis (Kertész, 1915), holotype, phallapodemeandpostgonite, overall, slightlysublateralview. 14–15 = phallapodemeand postgonites, ventral view: 14 = H. grandis (Kertész, 1915), holotype, 15 = H. pseudograndis sp.