Escaryus retusidens Attems, 1904

Nefediev, Pavel S., Farzalieva, Gyulli Sh., Tuf, Ivan H., Nedoev, Khozhiakbar Kh. & Niyazov, Saparmurad T., 2018, Millipede and centipede assemblages on the northern and southern slopes of the lowland Altais, southwestern Siberia, Russia (Diplopoda, Chilopoda), ZooKeys 741, pp. 219-254 : 235-237

Escaryus retusidens Attems, 1904

Escaryus retusidens - Titova 1972a: 110; 1972b: 135; Pereira and Hoffman 1993: 9; Volkova 2016: 675; Nefediev et al. 2017a: 11, 13: map; 2017c: 13; 2017d: 222: map.

Material examined

(all from Russia, southwestern Siberia, Altai Province, Charysh District, ca. 4.5 km SE of Charyshskoye Village). 1 ♂, 1 ♀ (ZMUM), 2 ♂♂, 4 juv. (ASU), Betula pendula and Populus tremula stand on N slope, 51°21'33.8"N, 83°37'23.2"E, 518 m a.s.l., 14.07.2015, leg P.N.; 1 ♂ (ASU), foot of S slope of mountain, Padus avium and Populus tremula stand near brook, hand sampling, 31.05.2016; 2 ♀♀, 3 juv. (ASU), site 1 on S slope, hand sampling, 31.05.2016; 2 juv. (ASU), site 1 on S slope, soil sample 1 (10-20 cm deep), 31.05.2016; 1 ♀ (ASU), site 1 on S slope, soil sample 1 (20-30 cm deep), 31.05.2016; 3 ♀♀ (ASU), site 1 on S slope, soil sample 3 (10-20 cm deep), 31.05.2016; 1 ♀, 3 juv. (ASU), site 1 on S slope, soil sample 3 (20-30 cm deep), 31.05.2016; 1 juv. (ASU), site 1 on S slope, soil sample 4 (20-30 cm deep), 1.06.2016; 1 juv. (ASU), site 1 on S slope, soil sample 5 (0-10 cm deep), 1.06.2016; 3 juv. (ASU), site 1 on S slope, soil sample 5 (10-20 cm deep), 1.06.2016; 1 ♂, 1 juv. (ASU), site 1 on S slope, soil sample 5 (20-30 cm deep), 1.06.2016; 1 ♂ (ASU), S slope between site 1 and site 2, broad gully with Padus avium , hand sampling, 1.06.2016; 1 juv. (ASU), site 2 on S slope, soil sample 1 (0-10 cm deep), 1.06.2016; 1 fragm. (ASU), site 2 on S slope, soil sample 1 (10-20 cm deep), 1.06.2016; 2 juv. (ASU), site 2 on S slope, soil sample 2 (0-10 cm deep), 1.06.2016; 1 juv. (ASU), site 2 on S slope, soil sample 2 (10-20 cm deep), 1.06.2016; 1 juv. (ASU), site 2 on S slope, soil sample 3 (0-10 cm deep), 1.06.2016; 1 juv. (ASU), site 2 on S slope, soil sample 5 (0-10 cm deep), 1.06.2016; 1 juv. (ASU), site 2 on S slope, hand sampling, 1.06.2016; 2 juv. (ASU), site 1 on N slope, soil sample 1 (0-10 cm deep), 2.06.2016; 2 ♀♀, 1 juv. (ASU), site 1 on N slope, soil sample 2 (0-10 cm deep), 2.06.2016; 2 juv. (ASU), site 1 on N slope, soil sample 2 (10-20 cm deep), 2.06.2016; 2 ♂♂, 2 ♀♀, 2 juv. (ASU), site 1 on N slope, soil sample 3 (0-10 cm deep), 2.06.2016; 1 ♂, 1 juv. (ASU), site 1 on N slope, soil sample 3 (10-20 cm deep), 2.06.2016; 1 ♀, 2 juv. (ASU), site 1 on N slope, soil sample 4 (0-10 cm deep), 2.06.2016; 1 ♂ (ASU), site 1 on N slope, soil sample 4 (10-20 cm deep), 2.06.2016; 1 ♂, 3 ♀♀ (ASU), site 1 on N slope, soil sample 5 (0-10 cm deep), 2.06.2016; 1 ♀ (ASU), site 1 on N slope, hand sampling, 2.06.2016; 1 ♀, 1 juv. (ASU), site 2 on N slope, soil sample 1 (0-10 cm deep), 2.06.2016; 3 juv. (ASU), site 2 on N slope, soil sample 2 (0-10 cm deep), 2.06.2016; 2 ♂♂, 2 ♀♀, 1 juv. (ASU), site 2 on N slope, soil sample 3 (0-10 cm deep), 2.06.2016; 1 ♀, 3 juv. (ASU), site 2 on N slope, soil sample 4 (0-10 cm deep), 2.06.2016; 1 juv. (ASU), site 2 on N slope, soil sample 4 (10-20 cm deep), 2.06.2016, all leg. P.N., Kh.N., S.N., V.S.; 1 ♀ (ASU), site 1 on N slope, hand sampling, 22.06.2016, leg. Kh.N.; 1 adult specimen (ASU), Betula pendula and Populus tremula stand on N slope, 51°21'33.8"N, 83°37'23.2"E, 518 m a.s.l., 12.07.2016, leg. P.N.; 1 juv. (ASU), site 1 on S slope, soil sample 1 (0-10 cm deep), 12.07.2016; 2 juv. (ASU), site 1 on S slope, soil sample 5 (10-20 cm deep), 12.07.2016; 1 ♂, 2 juv. (ASU), site 1 on N slope, soil sample 1 (10-20 cm deep), 13.07.2016; 1 fragm. (ASU), site 1 on N slope, soil sample 3 (0-10 cm deep), 13.07.2016; 2 ♀♀ (ASU), site 2 on N slope, soil sample 1 (0-10 cm deep), 13.07.2016; 1 juv. (ASU), site 2 on N slope, soil sample 2 (0-10 cm deep), 13.07.2016; 1 ♀, 1 juv. (ASU), site 2 on N slope, soil sample 4 (0-10 cm deep), 13.07.2016; 2 ♀♀ (ASU), site 2 on N slope, soil sample 4 (10-20 cm deep), 13.07.2016, all leg. Kh.N., S.N., V.S.; 1 juv., 2 fragm. (ASU), site 1 on S slope, soil sample 1 (10-20 cm deep), 22.08.2016; 1 fragm. (ASU), site 1 on S slope, soil sample 2 (10-20 cm deep), 22.08.2016; 1 ♀, 2 juv., 1 fragm. (ASU), site 1 on S slope, soil sample 4 (0-10 cm deep), 23.08.2016; 1 ♂ (ASU), site 2 on S slope, soil sample 5 (0-10 cm deep), 22.08.2016; 1 ♂, 2 ♀♀, 2 juv. (ASU), site 1 on N slope, soil sample 2 (0-10 cm deep), 23.08.2016; 1 juv. (ASU), site 1 on N slope, soil sample 4 (0-10 cm deep), 23.08.2016; 2 ♀♀, 1 juv. (ASU), site 1 on N slope, soil sample 5 (0-10 cm deep), 23.08.2016; 3 ♂♂, 1 ♀, 1 juv. (ASU), site 2 on N slope, soil sample 1 (0-10 cm deep), 23.08.2016; 1 ♀ (ASU), site 2 on N slope, soil sample 2 (0-10 cm deep), 23.08.2016; 2 ♀♀, 1 juv. (ASU), site 2 on N slope, soil sample 3 (0-10 cm deep), 23.08.2016; 1 ♀, 1 juv., 2 fragm. (ASU), site 2 on N slope, soil sample 3 (10-20 cm deep), 23.08.2016; 2 ♂♂, 1 ♀, 1 juv. (ASU), site 2 on N slope, soil sample 5 (0-10 cm deep), 23.08.2016; 1 ♀ (ASU), site 2 on N slope, hand sampling, 23.08.2016, all leg. P.N., Kh.N., S.N., V.S.; 3 ♂♂, 4 ♀♀, 3 juv. (ASU), Betula pendula and Populus tremula stand on N slope, 51°21'33.8"N, 83°37'23.2"E, 518 m a.s.l., 20.06.2017, leg. P.N.; 1 ♂ (ASU), site 1 on N slope, hand sampling, 23.06.2017, leg. P.N., Kh.N., A.A., E.A.

Distribution.

Central-Eastern-Palaearctic subboreal range: originally described from Kyrgyzstan, the species is widely distributed in Eurasia, spanning from the Black Sea region in the west through eastern Kazakhstan to Cisamuria in the east ( Titova 1972b). In Siberia E. retusidens has been known from the Kemerovo Area, Altai Province, and Republic of Altai ( Nefediev et al. 2017a, c, d).

Remarks.

In the study area, E. retusidens inhabits both slopes, and is one of the most dominant species.