Coenosia ignobilis Stein, 1911

Patitucci, Luciano Damian, Mulieri, Pablo Ricardo, Couri, Marcia Souto & Dominguez, Martha Cecilia, 2023, Phylogeny of the old and fragmented genus Austrocoenosia Malloch reveals new evidences on the morphology and evolution of the genera Coenosia Meigen and Neodexiopsis Malloch (Diptera: Muscidae), Arthropod Systematics & amp; Phylogeny 81, pp. 611-653 : 611

publication ID

publication LSID

persistent identifier

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Coenosia ignobilis Stein, 1911


Coenosia ignobilis Stein, 1911 View in CoL

Male (Fig. 17A).

Length. Body: 3.05-3.27 mm, wing: 3.00-3.10 mm. Head: (Fig. 17B View Figure 17 ). Frons at vertex about one third of the head width. Frons, fronto-orbital plate, parafacial and gena dark brown with grey pollinosity; 3-4 pairs of frontal setae. Frons longer than wide, with frontal triangle long, light grey, reaching lunula. Gena height less than the width of postpedicel. Fronto-orbital plate with 3-5 little setulae, close to parafacialia. Antenna black, apical angle of postpedicel acute; in lateral view inserted at the mid-level of the eye; arista with its longest microtrichia with similar length than its basal diameter. Thorax: (Fig. 17C View Figure 17 ). Black with grey pollinosity, with a dark-brown vitta along acrosticals setae and a vitta between dorsocentral and intra-alar rows of setae; anterior and posterior spiracles grey. Chaetotaxy: acr s short and irregular, the anterior presutural acr s with similar length to the eaDC; dorsocentrals 1+3, eaDC is less than one third of the aDC. Katepisternum with 2-3 setulae. Wing: Tegula black. Both calypters whitish hyaline with white margins; halter yellow. Legs: Black, apex of fore femur, basal half of mid and hind femora, and tibiae yellow; tarsus black. Fore femur with a row of strong pd, a row of strong pv setae, and a row of av setae at basal third. Mid femur with 5-6 setae on anterior surface on basal half, 2-3 pv and 2-3 av setae on basal third, 2 preapical setae on pd to posterior surface; mid tibia with one ad and one pd median setae, ad seta longer and placed more apically than pd seta. Hind femur with a complete row of ad and a row of av setae on apical half, with three pv seta on basal third, apical most is the longest of the three. Similar size of claws and pulvilli of the three legs. Abdomen: (Fig. 17D View Figure 17 ). Black, with grey pruinosity, tergites 3-5 with two dark brown spots. Tergite 6 partly visible in dorsal view. Sternite 5 broader than long, apical margin concave with a “U” shape, and with membrane; setae concentrated on the lobes and some long and strong on apical margin (Fig. 17E View Figure 17 ). Terminalia: Cercus curved in lateral view, keels only on basal two-thirds, apical margin straight (Fig. 17F, G View Figure 17 ). Surstylus longer than broad, wide at base and tapering towards apex, straight, sclerotized (Fig. 17H View Figure 17 ). Hypandrium tubular, wider than long, distal extreme exposing the phapod, and with several hairs on ventral surface. Aedeagus with pregt developed and sclerotized; pgt developed; epiphalus slightly sclerotized, and distiph tubular, sclerotized at base, and with spines on lateral surface (Fig. 17G, I-J View Figure 17 ).

Female (Fig. 18A, B).

Length. Body: 3.70-4.40 mm, wing: 3.50-4.24 mm. Differs from male as follows: Legs: Mid and hind femora more yellow, hind tibia with one long ad, one short av, and one long pd setae on middle third. Ovipositor: Tergites 6, 7, and 8 with two long and wide sclerotized plates; epiproct triangular, with hair-like setae, shorter than cercus (Fig. 18C View Figure 18 ). Sternites 6 and 7 with one long and wide plate; sternite 8 with one central plate and two small, linear, and sclerotized plates each with 4-5 setae on distal margin; hypoproct triangular, setulose, with several strong setae on distal margin (Fig. 18D View Figure 18 ). Spermathecae (Fig. 18E View Figure 18 ).

Type material.

Paratype. Male; ZMUH [pinned, in good condition]. Original labels: "PARALECTOTYPE ♂ / Coenosia / Coenosia ignobilis / Stein, 1911 / Des.A.C. Pont 1999" printed, on white paper; " Coenosia / Coenosia ignobilis / Typo Stein" handwritten, on white paper; “Type” printed on red paper; "Chile [printed] / 20.IX.02 [handwritte] / Quillota [printed]" on green paper.

Additional material examined.

ARGENTINA - Chubut province • 2 females; PNLA, Cabana La Cascada ; -42.888499, -71.592376; 532 m a.s.l.; Feb 2013; Mulieri, Patitucci & Olea leg.; MACN GoogleMaps 5 females, 5 males; PNLA, Seccional Lago Verde ; -42.718803, -71.727470; 538 m a.s.l.; Oct 2014; Mulieri & Patitucci leg.; MACN GoogleMaps 1 female; PNLA, Puerto Mermoud ; -42.723248; -71.748801; 521 m a.s.l.; 25 Oct 2014; Torretta leg.; MACN GoogleMaps 3 males; PNLP, Gendarmería; -42.097468, -71.681953; 200 m a.s.l.; 07-11 Jan 2012; Mulieri & Patitucci leg.; MACN GoogleMaps 1 male; PNLP, Intendencia ; -42.085077, -71.614662; 195 m a.s.l.; 16 Jan 2011; Mulieri & Patitucci leg.; MACN GoogleMaps 1 female, 9 males; PNLP, La Playita ; -42.099032, -71.607425; 205 m a.s.l.; 09 Jan 2012; Patitucci leg.; MACN GoogleMaps 1 female; PNLP, Rio Azul ; -42.0916, -71.6155; 184 m a.s.l.; 17 Jan 2011; Mulieri & Patitucci leg.; MACN GoogleMaps 2 females; PNLP, Rio Turbio ; -42.228541, -71.666482; 204 m a.s.l.; 12 Jan 2011; Mulieri & Patitucci leg.; MACN GoogleMaps . - Neuquén province • 1 female; Cerro Chapelco ; -40.197050, -71.298453; 1400 m a.s.l.; 15 Feb 1983; M. Gentili & P. Gentili leg.; IFML GoogleMaps 4 females, 1 male; San Martín de los Andes, Truran Kura ; 1000 m a.s.l.; 27 Sep 1986; M. Gentili & P. Gentili leg.; IFML . - Río Negro province • 3 males; PNNH, Lago Guillelmo ; -41.358807, -71.515071; 833 m a.s.l.; 14 Sep-01 Oct 2008; Garré & Montes de Oca leg.; MLP; Malaise GoogleMaps . CHILE - Región de la Araucanía • 1 female; Angol ; -37.7800, -72.7500; 15 Sep 1926; JR Malloch det.; SM Lopes leg. MNRJ GoogleMaps .

Distribution (Fig. 27G).

ARGENTINA: Chubut (new record), Neuquén (new record), Río Negro. CHILE: Arch. Juan Fernandez, R. de la Aruacania, R. de Valparaiso.


Stein (1911) described Coenosia ignobilis with male and female specimens from Chile. Later, Malloch (1934) studied a large series of male and female specimens from Argentina and Chile, presented a brief description, and considered it as part of Austrocoenosia . Later, Couri and Albuquerque (1979) observed a female specimen from the series of specimens studied by Malloch (op. cit), and included the species as part of the genus Neodexiopsis . Pont (2001) designated lectotypes and paralectotypes and established the original combination Coenosia ignobilis . Recently, Couri and Pont (2020) presented a redescription of the paralectotype housed in ZMUH.