Archaeohelorus tensus, Shi, Xiaoqing, Zhao, Yunyun, Shih, Chungkun & Ren, Dong, 2014

Shi, Xiaoqing, Zhao, Yunyun, Shih, Chungkun & Ren, Dong, 2014, Two new species of Archaeohelorus (Hymenoptera, Proctotrupoidea, Heloridae) from the Middle Jurassic of China, ZooKeys 369, pp. 49-59 : 53-55

Archaeohelorus tensus sp. n. Figure 3

Etymology.

The specific name “tensus” means stretching and long, referring to the shape of the forewing.

Type material.

Holotype, CNU-HYM-NN2012056p/c, part and counterpart, dorsal view, male. A well-preserved almost complete body with antenna, forewings and part of legs and partial hind wings.

Locality and age.

Jiulongshan Formation, Middle Jurassic, Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.

Diagnosis.

Forewing 2cu-a postfurcal with intersection of Cu and Rs+M. First abscissa of Rs (1-Rs) and basal section of M (1-M) arched toward basal of wing. Hind wing with tubular veins C, Sc+R, and Rs preserved.

Description.

A smalladult specimen with a total body length of 4.4 mm. Head suboval and large relative to mesosoma in width. Eyes large, and located at the both sides of the head.

Antennae filiform, thick, with 6 segments preserved on left and 5 segments preserved on right. Scape swollen and bell-shaped; pedicel short and quadrate; subsequent flagellomeres not well preserved.

Mesosoma subhexagonal and broader than head; pronotum narrower than head, short, probably covered by mesonotum; mesoscutum distorted in the middle and trapezoidal with notauli not distinct but preserved; tegula large and subtriangular; scutellum round with pits anteriorly; propodeum narrow, transverse and areolated.

Metasoma suboval with seven segments; first metasomal segment transverse with several longitudinal ridges; second segment trapezoidal with less longitudinal ridges anteriorly; other ones smooth; the male terminalia triangular, partially covered by the previous segment.

Forewing broad and subtriangular. Pterostigma long and acute apically, not widened beyond mid-length. C robust and extending near the apex of forewing. 2r-rs arising from basal one third of pterostigma, slightly longer than the width of pterostigma, and slightly oblique apicad. R robust and cell r closed with C, R and 1-RS. Rs straight and intersecting with distal part of C. Cell 1+2r longer and narrower relatively and six-sided surrounded by R, 1-Rs, 1-Rs+M, 2-Rs+M, 2-Rs, and 2r-rs. M+Cu straight and distinct; M and Cu distinct, almost straight. 1-Rs as long as 1-M and slightly inclined toward wing base. Cell 1mcu small, subtriangular and relatively slender with 2-M+Cu 4.3 times as long as 1-M. M and Rs branching at 30% from 1m-cu of the length between 1m-cu and 2r-rs. 1cu-a and 2cu-a distinct and reaching A. 1cu-a in line with 1-M; and 2cu-a postfurcal with Rs+M distinctly.

Hind wing, with tubular veins C parallel with Sc+R at base, Rs short, intersecting distal part of C.

Left foreleg, midleg and hindleg and right foreleg with only coxa and femur partially preserved, hind femur much thicker, nearly three times as long as wide; right midleg with partial spindle-shaped femur and partial tibia preserved, right hindleg with partial coxa, trapezoidal trochanter, robust spindle-shaped femur and relatively thin tibia and some parts of tarsus preserved.

Measurements (in mm): Body length 4.67, head length1.39, width 0.68, mesosoma length 1.68, width 1.36; metasoma 2.23 long; lengths of metasomal segments are 0.20, 0.39, 0.48, 0.38, 0.25 and 0.32; forewing length> 2.96, width> 1.36.

Remarks and comparison.

This species is assigned to Archaeohelorus Shih, Feng & Ren, 2011 by its seven separated metasomal segments, the first metasomal segment transverse with several longitudinal ridges. Forewing 1-Rs as long as 1-M. Cell 1mcu small and subtriangular, 1cu-a in line with 1-M. Compared with Archaeohelorus hoi , Archaeohelorus tensus has preserved hind wing with tubular veins C, Sc+R, R, and Rs preserved; in forewing, 2cu-a distinctly postfurcal with intersection of Cu and Rs+M (vs. 2cu-a intersecting Cu and Rs+M at the same point in Archaeohelorus hoi ); cell 1mcu distinctly slender, 4.3 times as long as wide (vs. 2.4 times in Archaeohelorus hoi ); 1-Rs and 1-M arched toward basal of wing (vs. 1-Rs and 1-M straight in Archaeohelorus hoi ). It also differs from Archaeohelorus polyneurus by its postfurcal 2cu-a, slender cell 1mcu.